Cystopteris protrusa

Family:CYSTOPTERIDACEAE
Species:Cystopteris protrusa (Weatherby) Blasdell
Common Name:Spreading Bladder Fern; Lowland Bladder Fern
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:**
Plant Notes:**
Taxonomic Notes:**
Status:Native
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyCYSTOPTERIDACEAE - Brittle Fern Family family
Genus Cystopteris
Species Cystopteris protrusa (Weatherby) Blasdell - Spreading Bladder Fern; Lowland Bladder Fern

Citation

Citation CYSTOPTERIS PROTRUSA (Weatherby) Blasdell, Mem. Torry Bot. Club 21(4): 41. 1963.
Basionym: Cystopteris fragilis (Linnaeus) Bernhardi, var. protrusa Weatherby 1935.
Type: TENNESSEE: Knox Co.: Knoxville, 5 Jun 1898, Ruth 564 (holotype: GH).

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Cystopteris fragilis var. protrusa Cystopteris fragilis (Linnaeus) Bernhardi, var. protrusa Weatherby, Rhodora 37: 373, t. 383(1, 5). 1935. TENNESSEE: Knox Co.: Knoxville, 5 Jun 1898, Ruth 564 (holotype: GH).

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Bibb, Blount, Calhoun, Cherokee, Cleburne, Colbert, Etowah, Fayette, Jackson, Jefferson, Lauderdale, Lawrence, Limestone, Madison, Marion, Marshall, Morgan, Shelby, St. Clair, Talladega, Tallapoosa, Tuscaloosa, Walker, Winston

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AMAL, AUA, JSU, TROY, UNA, UNAF, UWAL

Range of years during which specimens were collected: 1941 - 2015

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
AMAL00013294 Colbert 17 Apr 2006 Keener, Brian R.
2800
AMAL
Cystopteris protrusa (Weatherby) Blasdell
UNA00052666 Madison 19 May 1980 Meigs, Russell A.
532
UNA
Cystopteris protrusa (Weatherby) Blasdell
UNA00047564 Madison 20 Apr 1973 Simmers, Richard W.
2724
UNA
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001174 Colbert 15 Jun 1991 Stewart, Donald
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106667 Blount 27 Jun 1998 Keener, Brian R.
1099
JSU
Cystopteris protrusa (Weatherby) Blasdell
UWAL0037847 Calhoun 14 Jul 2017 Spaulding, Daniel D.
14826
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106686 Morgan 13 Apr 1997 Spaulding, Daniel D.
9729
JSU
Cystopteris protrusa (Weatherby) Blasdell
AUA_ACC_31791 Colbert 28 Jun 1976 Short, John W.
713
AUA
Cystopteris Protrusa (Weatherby) Blasdell
UNAF0001268 Colbert 10 Oct 1989 Hammond, Kathryn
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106680 Jackson 04 Jun 1975 Whetstone, R. David
4269A
JSU
Cystopteris protrusa (Weatherby) Blasdell
UWAL0033907 Fayette 30 Mar 2016 Keener, Brian R.
9443
UWAL
Cystopteris protrusa (Weatherby) Blasdell
UWAL0036961 Lawrence 07 Jun 2017 England, J. Kevin
7921
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106687 Morgan 29 Mar 1997 Whetstone, R. David
s.n.
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00025740 Colbert 29 Jun 1941 Shaw, James W.
s.n.
UNA
Cystopteris fragilis (L.) Bernh. var. protrusa Weatherby
UNAF0001177 Marion 28 Oct 1989 Spiller, Debra
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106675 Jackson 29 Apr 1998 Brodeur, Kristin R.
582
JSU
Cystopteris protrusa (Weatherby) Blasdell
UWAL0044779 Jackson 02 Jun 2018 England, J. Kevin
9213
UWAL
Cystopteris protrusa (Weatherby) Blasdell
AUA_ACC_31792 Morgan 19 Mar 1977 Short, John W.
876
AUA
Cystopteris Protrusa (Weatherby) Blasdell
UNAF0001176 Marion 28 Oct 1989 Crews, Sherry
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106679 Jackson 01 Jun 1993 Whetstone, R. David
s.n.
JSU
Cystopteris protrusa (Weatherby) Blasdell
UWAL0050817 Jefferson 05 Oct 1965 Kinnear, Sue
Steeves 32-1
UWAL
Cystopteris protrusa
UWAL0047997 Cherokee 27 Mar 2020 England, J. Kevin
10829
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106672 Etowah 06 Apr 1995 Hodge, Lesley M.
4122
JSU
Cystopteris protrusa (Weatherby) Blasdell
UWAL0001198 St. Clair 02 Jun 2008 Keener, Brian R.
4556
UWAL
Cystopteris protrusa (Weatherby.) Blasdell
UNAF0001175 Colbert 07 Sep 1984 Moir, Judith
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
AUA_ACC_35187 Cleburne 3 Jun 1978 Glick, J.I.
AUA
Cystopteris Protrusa (Weatherby) Blasdell
AUA_ACC_71934 Tuscaloosa 16 Jun 1983 Short, John W.
1327
AUA
Cystopteris Protrusa (Weatherby) Blasdell
UNAF0001170 Colbert 08 Sep 1985 Cantrell, Lanetta
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106682 Limestone 01 Mar 1997 Hofmann, Tim L.
00303
JSU
Cystopteris protrusa (Weatherby) Blasdell
ALNHS00003743 Lauderdale 29 Sep 2015 Barger, T. Wayne
SC-513
ALNHS
Cystopteris protrusa (Weatherby) Blasdell
UWAL0052502 Etowah 10 May 1970 Dean, Blanche E.
237
UWAL
Cystopteris protrusa
JSU106678 Jackson 28 Mar 1976 Atkinson, Tim A.
965
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052668 Morgan 24 May 1980 Meigs, Russell A.
636
UNA
Cystopteris protrusa (Weatherby) Blasdell
UWAL0031316 Lauderdale 17 Apr 2011 England, J. Kevin
2794
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106676 Jackson 04 May 1994 Spaulding, Daniel D.
6071
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052671 Tuscaloosa 27 Apr 1874 Smith, Eugene A.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
AMAL00014385 St. Clair 02 Jun 2008 Keener, Brian R.
4556
AMAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106692 Walker 13 Apr 1970 Kral, Robert
38396
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052672 Colbert 12 Mar 1953 Chermock, Ralph L.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106690 Talladega 18 Mar 1995 Spaulding, Daniel D.
8891
JSU
Cystopteris protrusa (Weatherby) Blasdell
AUA000039560 Limestone 9 Apr 2016 England, J. Kevin
6179
AUA
Cystopteris Protrusa (Weatherby) Blasdell
JSU106673 Etowah 19 Mar 1995 Hodge, Lesley M.
4092
JSU
Cystopteris protrusa (Weatherby) Blasdell
TROY000013369 Colbert 17 Apr 2006 Keener, Brian R.
2800
TROY
Cystopteris protrusa (Weatherby) Blasdell
UNA00052674 Tuscaloosa 11 Jun 1938 Stevenson
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106674 Etowah 21 May 1976 Atkinson, Tim A.
1505
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052667 Madison 23 May 1980 Meigs, Russell A.
624
UNA
Cystopteris protrusa (Weatherby) Blasdell
UWAL0001197 Colbert 17 Apr 2006 Keener, Brian R.
2800
UWAL
Cystopteris protrusa (Weatherby.) Blasdell
UNA00052670 Tuscaloosa 12 Jun 1877 Smith, Eugene A.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001172 Colbert 07 Sep 1984 Thompson, Jennifer
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106681 Jefferson 05 Oct 1965 Kinnear, Sue
32
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00058266 Blount 02 May 1953 Chermock, Ralph L.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106685 Marshall 13 Mar 1992 Spaulding, Daniel D.
1169
JSU
Cystopteris protrusa (Weatherby) Blasdell
AUA_ACC_31793 Morgan 5 Jul 1977 Short, John W.
911
AUA
Cystopteris Protrusa (Weatherby) Blasdell
JSU106669 Etowah 12 Jun 1994 Hodge, Lesley M.
2363
JSU
Cystopteris protrusa (Weatherby) Blasdell
JSU106666 Bibb 31 Jul 1998 Whetstone, R. David
19656
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052675 Madison _ Jul 1954 Brown, Jack S.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106668 Cleburne 15 Oct 1996 Hruska, Susan E.
1456
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052673 Morgan 08 Mar 1953 White, Jesse S.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
UWAL0036433 Marshall 13 Aug 2017 England, J. Kevin
8411
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106677 Jackson 04 May 1994 Spaulding, Daniel D.
6120
JSU
Cystopteris protrusa (Weatherby) Blasdell
TROY000028131 St. Clair 02 Jun 2008 Keener, Brian R.
4556
TROY
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001169 Colbert 28 Nov 1984 Hardy, Rhonda
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
UWAL0032385 Limestone 09 Apr 2016 England, J. Kevin
6179
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106693 Winston 15 Oct 1975 Radford, Albert E.
s.n.
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00052676 Tuscaloosa 19 Apr 1953 Chermock, Ralph L.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106684 Limestone 25 Apr 1996 Whetstone, R. David
17428
JSU
Cystopteris protrusa (Weatherby) Blasdell
UWAL0015451 Colbert 15 Apr 2013 Spaulding, Daniel D.
13719
UWAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106670 Etowah 26 Apr 1994 Hodge, Lesley M.
1236
JSU
Cystopteris protrusa (Weatherby) Blasdell
AMAL00005355 Madison 27 Jul 1996 Duke, J.F.
774
AMAL
Cystopteris protrusa (Weatherby) Blasdell
UWAL0031337 Limestone 27 Apr 2012 England, J. Kevin
3359
UWAL
Cystopteris protrusa (Weatherby) Blasdell
UNA00052669 Tuscaloosa 19 May 1979 Wiersema, John H.
719B
UNA
Cystopteris protrusa (Weatherby) Blasdell
AMAL00019762 Tuscaloosa 16 Jun 1983 Short, John W. (Jack)
1327
AMAL
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001252 Marion 20 Oct 1989 Lolley, Kim
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
ALNHS00007152 Colbert 11 Jun 2021 Barger, T. Wayne
SP#5542
ALNHS
Cystopteris protrusa (Weatherby) Blasdell
AMAL00002495 Colbert 15 Apr 2013 Spaulding, Daniel D.
13719
AMAL
Cystopteris protrusa (Weatherby) Blasdell
ALNHS00003444 Tallapoosa 29 Apr 2015 Barger, T. Wayne
SP#1686
ALNHS
Cystopteris protrusa (Weatherby) Blasdell
AUA_ACC_55689 Bibb 1 Apr 1972 Kral, Robert
45425
AUA
Cystopteris Protrusa (Weatherby) Blasdell
AMAL00023552 Colbert 23 Aug 2015 Spaulding, Daniel D.
14386
AMAL
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001171 Colbert 15 Oct 1983 Bobo, Wade
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
JSU106683 Limestone 01 Apr 1997 Hofmann, Tim L.
00359
JSU
Cystopteris protrusa (Weatherby) Blasdell
ALNHS00004000 Colbert 12 Apr 2016 Barger, T. Wayne
SP#2193
ALNHS
Cystopteris protrusa (Weatherby) Blasdell
JSU106689 Talladega 10 Apr 1981 Whetstone, R. David
10511
JSU
Cystopteris protrusa (Weatherby) Blasdell
TROY000031035 Morgan 05 Jul 1977 Short, John W. (Jack)
911
TROY
Cystopteris protrusa (Weatherby) Blasdell
UNA00058265 Blount 21 Mar 1953 White, Jesse S.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106688 Shelby 11 Oct 1997 Spaulding, Daniel D.
10166
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001173 Colbert 21 Oct 1986 Henderson, Cathy
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
UNA00064787 Colbert 17 Apr 2006 Keener, Brian R.
2800
UNA
Cystopteris protrusa (Weatherby) Blasdell
JSU106691 Tuscaloosa 16 Oct 1975 Whetstone, R. David
7218
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00053201 Tuscaloosa 19 Apr 1953 Chermock, Ralph L.
s.n.
UNA
Cystopteris protrusa (Weatherby) Blasdell
TROY000050823 Lauderdale 17 Apr 2011 England, J. Kevin
2794
TROY
Cystopteris protrusa (Weatherby) Blasdell
AMAL00023143 Lauderdale 17 Apr 2011 England, Kevin J.
2794
AMAL
Cystopteris protrusa (Weatherby) Blasdell
JSU106671 Etowah 27 May 1994 Hodge, Lesley M.
1351
JSU
Cystopteris protrusa (Weatherby) Blasdell
UNA00054398 Blount 27 Jun 1998 Keener, Brian R.
1099
UNA
Cystopteris protrusa (Weatherby) Blasdell
UNAF0001269 Lauderdale 26 Oct 1989 Wieseman, Kevin
s.n.
UNAF
Cystopteris protrusa (Weatherby) Blasdell
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