Thelypteris palustris var. pubescens

Family:THELYPTERIDACEAE
Species:Thelypteris palustris Schott var. pubescens (Lawson) Fernald
Common Name:Eastern Marsh Fern
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:THPAP
Plant Notes:Eastern Marsh Fern is a native herbaceous perennial in the Marsh Fern family (Thelypteridaceae). In occurs throughout Alabama. Eastern Marsh Fern occurs in low woods, along creeks, in wet roadside ditches, and in low pastures. It is a perennial with a long creeping rhizome. The leaves (fronds) are erect to arching, elliptic to lanceolate in outline, broadest near the middle, and once pinnate with lobed pinna. The leaves are deciduous. The petiole is yellow-green in color, about as long as the blade, and brittle. The pinna lobes are rounded to slightly pointed at the apex, and the basal veins from adjacent pinna lobes join before reaching the sines. The pinna are lobed nearly to their base. The veins are forked. The leaf is slightly pubescent on the veins, but the blade tissue is glabrous. Eastern Marsh Fern reproduces by forming spores. The spores are produced in sporangia that are grouped together in clusters known as sori. The sori are located about mid-way between the main vein of the pinna lobes and the margin. The sori are covered with a round, pubescent flap of tissue known as the indusium. Eastern Marsh Fern often occurs in saturated soils or even shallow water in full sunlight. It is sometimes available from nurseries. It requires a constantly moist soil and is one of the few ferns that thrive in direct sun. Eastern Marsh Fern is a vigorous grower wide spreading slender rhizomes. Under the right conditions, and can become “weedy” in the garden.--A. Diamond
Taxonomic Notes:**
Status:Native
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyTHELYPTERIDACEAE - Marsh Fern family
Genus Thelypteris
Species Thelypteris palustris Schott var. pubescens (Lawson) Fernald - Eastern Marsh Fern

Citation

Citation THELYPTERIS PALUSTRIS Schott, var. PUBESCENS (G. Lawson) Fernald, Rhodora 31: 34. 1929.
Basionym: Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864.
Type: CANADA: Ontario: Addington Co.: Odessa, s.n., Lawson s.n. (holotype: ?).

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Dryopteris thelypteris subsp. pubescens Dryopteris thelypteris (Linnaeus) A. Gray, subsp. pubescens (G. Lawson) Hultén, Kongl. Svenska Vetenskapsakad. Handl. 8(5): 180. 1962.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Dryopteris thelypteris var. haleana Dryopteris thelypteris (Linnaeus) A. Gray, var. haleana (Fernald) M. Broun ex Weatherby, Amer. Fern J. 26: 95. 1936.Thelypteris palustris Schott, var. haleana Fernald 1929. 
Dryopteris thelypteris var. kamschatica Dryopteris thelypteris (Linnaeus) A. Gray, var. kamschatica C. Christensen, in Hultén, Fl. Kamtchat. 1: 38. 1927.  
Dryopteris tremula Dryopteris tremula Christ, Notul. Syst. (Paris) 1: 234. 1910. MEXICO: Michoacán: La Huerta, vicinity of Morelia, 15 Oct 1909, Arsène 3106 (holotype; P?; isotypes: F, GH, MEXU(3 shts), MO, NY, U).
Dryopteris thelypteris var. pubescens Dryopteris thelypteris (Linnaeus) A. Gray, var. pubescens (G. Lawson) Nakai, Bot. Mag. (Tokyo) 45: 97. 1931; (G. Lawson) Komarov, Izv. Bot. Sada Akad. Nauk SSSR 30: 193. 1932.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Filix-mas thelypteris var. pubescens Filix-mas thelypteris (Linnaeus) Farwell, var. pubescens (G. Lawson) Farwell, Amer. Midl. Naturalist 12: 258. 1931.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Lastrea thelypteris var. pubescens Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson, Edinburgh New Philos. J., ser. 2. 19: 277. 1864. CANADA: Ontario: Addington Co.: Odessa, s.n., Lawson s.n. (holotype: ?).
Thelypteris confluens var. pubescens Thelypteris confluens (Thunberg) C. V. Morton, var. pubescens (G. Lawson) Pringle, Techn. Bull. Roy. Bot. Gard. Hamilton 4: 42. 1969.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Thelypteris palustris subsp. pubescens Thelypteris palustris Schott, subsp. pubescens (G. Lawson) Holub, Folia Geobot. Phytotax. 4: 45. 1969.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Thelypteris palustris var. haleana Thelypteris palustris Schott, var. haleana Fernald, Rhodora 31: 34, f. 5-6. 1929. LOUISIANA: Rapides Par.: Alexandria, s.d., Hale s.n. (holotype: GH).
Thelypteris thelypteris    

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Bibb, Butler, Choctaw, Clarke, Clay, Coffee, Conecuh, Covington, Crenshaw, Elmore, Escambia, Etowah, Lawrence, Lee, Limestone, Marengo, Mobile, Montgomery, Perry, Pike, Talladega, Tallapoosa, Tuscaloosa

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: AMAL, AUA, JSU, TROY, UNA, USAM, UWAL

Range of years during which specimens were collected: -

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
TROY000043289 Lawrence 08 Jun 2014 J. Kevin England
4918
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
JSU109756 Clarke 19 Jul 1970 R. Kral
40191
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
63823 Escambia 17 JUN 2004 DIAMOND, JR., ALVIN R.
14442
AUA
THELYPTERIS PALUSTRIS SCHOTT
TROY000029603 Baldwin 16 Jun 2009 Terry Wayne Barger
623
TROY
Thelypteris palustris Schott
TROY000000478 Crenshaw 03 Jun 2001 Alvin R. Diamond
12470
TROY
Thelypteris palustris Schott
UNA00052241 Tuscaloosa 11 May 1953 R. L. Chermock
s.n.
UNA
Thelypteris palustris Schott
AMAL00019903 Limestone 26 Jul 1983 John W. Short
1364
AMAL
Thelypteris palustris (A. Gray) Schott
31842 Autauga 20 JUL 1976 SHORT, JOHN W.
772
AUA
THELYPTERIS PALUSTRIS SCHOTT
UWAL0017235 Lawrence 08 Jun 2014 England, J. Kevin
4918
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000030804 Autauga 20 Jul 1976 Jack W. Short
772
TROY
Thelypteris palustris Schott
USAM000000294 Baldwin 29 Jun 1974 Michel G. Lelong
7992
USAM
Thelypteris palustris Schott
UNA00068858 Perry 7 Aug 2007 Brian R. Keener
3836
UNA
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UNA00068909 Baldwin 6 Jun 2006 D. D. Spaulding
12467
UNA
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UWAL0002857 Baldwin 09 Sep 2010 Diamond, A.R.
21560
UWAL
Thelypteris palustris Schott
AMAL00009615 Baldwin 21 Apr 2011 T. Wayne Barger
311
AMAL
Thelypteris palustris Schott
UNA00052240 Choctaw 5 Aug 1953 R. L. Chermock
s.n.
UNA
Thelypteris palustris Schott
AMAL00012385 Choctaw 03 Jul 1980 R. Kral
65913
AMAL
Thelypteris palustris Schott
31769 Lee 21 APR 1977 SHORT, JOHN W.
890
AUA
THELYPTERIS PALUSTRIS SCHOTT
UWAL0030070 Montgomery 10 Aug 2015 Diamond, Alvin R.
26745
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UNA00052253 Lee 21 Apr 1977 John W. Short
890
UNA
Thelypteris palustris Schott
AMAL00017584 Escambia 17 Jun 2004 A.R. Diamond
14442
AMAL
Thelypteris palustris Schott
JSU109758 Clay 9 Oct 1982 R. D. Whetstone
12715
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000013477 Tallapoosa 11 Apr 2003 Daniel D. Spaulding
11718
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UWAL0016925 Baldwin 13 Sep 2008 England, J. Kevin
1093
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UNA00052245 Autauga 28 May 1953 Jesse S. White
s.n.
UNA
Thelypteris palustris Schott
JSU109759 Etowah 16 Sep 1995 L. M. Hodge-Spaulding
4352
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AMAL00019767 Baldwin 04 Aug 1982 John W. Short
1262
AMAL
Thelypteris palustris (A. Gray) Schott
31773 Baldwin 2 JUN 1976 SHORT, JOHN W.
601
AUA
THELYPTERIS PALUSTRIS SCHOTT
TROY000000476 Conecuh 26 Aug 2000 Alvin R. Diamond
12014
TROY
Thelypteris palustris Schott
UWAL0001419 Perry 07 Aug 2007 Keener, Brian R.
3836
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
55130 Pike 7 AUG 1995 DIAMOND, JR., ALVIN R.
9824
AUA
THELYPTERIS PALUSTRIS SCHOTT
UWAL0030421 Bibb 07 Aug 2015 England, J. Kevin
5570
UWAL
Thelypteris palustris (Schott) var. pubescens (Lawson) Fernald
TROY000033638 Barbour 01 Oct 2011 Derek Morrison
129
TROY
Thelypteris palustris Schott
UNA00056167 Mobile 5 Jun 1953 Jack S. Brown
s.n.
UNA
Thelypteris palustris Schott
TROY000000479 Escambia 17 Jun 2004 Alvin R. Diamond
14442
TROY
Thelypteris palustris Schott
UWAL0012711 Marengo 25 Jul 2011 England, J. Kevin
3097
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
31770 Escambia 16 SEP 1976 SHORT, JOHN W.
831
AUA
THELYPTERIS PALUSTRIS SCHOTT
TROY000032980 Baldwin 09 Sep 2010 Alvin R. Diamond
21560
TROY
Thelypteris palustris Schott
72231 Lawrence 8 JUN 2014 ENGLAND, J. KEVIN
4918
AUA
THELYPTERIS PALUSTRIS SCHOTT var. PUBESCENS (LAWSON) FERN.
TROY000013301 Baldwin 06 Jun 2006 Daniel D. Spaulding
12467
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UNA00052252 Autauga 20 Jul 1976 John W. Short
772
UNA
Thelypteris palustris Schott
AMAL00005512 Pike 11 Jul 2001 A.R. Diamond
12523
AMAL
Thelypteris palustris Schott
UWAL0001418 Baldwin 06 Jun 2006 Spaulding, Daniel D.
12467
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UNA00057505 Clarke 25 Aug 1953 Jesse S. White
s.n.
UNA
Thelypteris palustris Schott
AMAL00011934 Coffee 14 Nov 1978 R. Kral
63129
AMAL
Thelypteris palustris Schott
AMAL00005511 Mobile 05 Jun 1953 J.S. Brown
s.n.
AMAL
Thelypteris palustris Schott
31768 Mobile 2 JUN 1976 SHORT, JOHN W.
590
AUA
THELYPTERIS PALUSTRIS SCHOTT
TROY000031556 Butler 15 Jul 2010 Alvin R. Diamond
21462
TROY
Thelypteris palustris Schott
USAM000000641 Lawrence 4 Jul 2003 R. Pate
72
USAM
Thelypteris palustris Schott
TROY000000475 Coffee 14 Nov 1978 Robert Kral
63129
TROY
Thelypteris palustris Schott
UNA00052246 Mobile 22 May 1894 Charles Mohr
s.n.
UNA
Thelypteris palustris Schott
UNA00052239 Talladega s.d. J. C. O'Kelley
s.n.
UNA
Thelypteris palustris Schott
TROY000000480 Pike 11 Jul 2001 Alvin R. Diamond
12523
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000042710 Montgomery 10 Aug 2015 Alvin R. Diamond
26745
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
31841 Conecuh 16 SEP 1976 SHORT, JOHN W.
826
AUA
THELYPTERIS PALUSTRIS SCHOTT
34605 Clarke 8 MAY 1976 SHORT, JOHN W.
506
AUA
THELYPTERIS PALUSTRIS SCHOTT
AMAL00014169 Perry 07 Aug 2007 Brian R. Keener
3836
AMAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
TROY000043989 Bibb 07 Aug 2015 J. Kevin England
5570
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
JSU109757 Elmore 1 Aug 1978 R. Kral
62363
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AMAL00013561 Baldwin 06 Jun 2006 Daniel D. Spaulding
12467
AMAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UWAL0030596 Covington 15 Aug 1987 Kral, R.
74303
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UNA00052244 Autauga 28 May 1953 Jesse S. White
s.n.
UNA
Thelypteris palustris Schott
UNA00063078 Covington 17 Jun 2004 Alvin R. Diamond
14442
UNA
Thelypteris palustris Schott
JSU109760 Etowah 16 Sep 1995 L. M. Hodge-Spaulding
4352
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AMAL00019891 Baldwin 04 Aug 1982 John W. Short
1266
AMAL
Thelypteris palustris (A. Gray) Schott
31771 Covington 2 JUN 1976 SHORT, JOHN W.
623
AUA
THELYPTERIS PALUSTRIS SCHOTT
TROY000000477 Crenshaw 27 Jul 2000 Alvin R. Diamond
11928
TROY
Thelypteris palustris Schott
TROY000000474 Choctaw 03 Jul 1980 Robert Kral
65913
TROY
Thelypteris palustris Schott
USAM000000295 Mobile 6 Sep 1978 Michel G. Lelong
10504
USAM
Thelypteris palustris Schott
UNA00052247 Montgomery _ Oct 1879 Charles Mohr
s.n.
UNA
Thelypteris palustris Schott
UNA00052248 Baldwin 28 Apr 1947 Francis X. Lueth
L 24
UNA
Thelypteris palustris Schott
UWAL0002113 Butler 15 Jul 2010 Diamond, A.R.
21462
UWAL
Thelypteris palustris Schott
AMAL00020272 Baldwin 21 Apr 2011 T. Wayne Barger
SP#311
AMAL
Thelypteris palustris Schott
TROY000040562 Marengo 25 Jul 2011 J. Kevin England
3097
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
31772 Butler 15 SEP 1976 SHORT, JOHN W.
819
AUA
THELYPTERIS PALUSTRIS SCHOTT
71968 Baldwin 4 AUG 1982 SHORT, JOHN W.
1262
AUA
THELYPTERIS PALUSTRIS SCHOTT
TROY000014447 Perry 07 Aug 2007 Brian R. Keener
3836
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
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