Thelypteris palustris var. pubescens

Family:THELYPTERIDACEAE
Species:Thelypteris palustris Schott var. pubescens (G. Lawson) Fernald
Common Name:Eastern Marsh Fern
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:THPAP
Plant Notes:Eastern Marsh Fern is a native herbaceous perennial in the Marsh Fern family (Thelypteridaceae). In occurs throughout Alabama. Eastern Marsh Fern occurs in low woods, along creeks, in wet roadside ditches, and in low pastures. It is a perennial with a long creeping rhizome. The leaves (fronds) are erect to arching, elliptic to lanceolate in outline, broadest near the middle, and once pinnate with lobed pinna. The leaves are deciduous. The petiole is yellow-green in color, about as long as the blade, and brittle. The pinna lobes are rounded to slightly pointed at the apex, and the basal veins from adjacent pinna lobes join before reaching the sines. The pinna are lobed nearly to their base. The veins are forked. The leaf is slightly pubescent on the veins, but the blade tissue is glabrous. Eastern Marsh Fern reproduces by forming spores. The spores are produced in sporangia that are grouped together in clusters known as sori. The sori are located about mid-way between the main vein of the pinna lobes and the margin. The sori are covered with a round, pubescent flap of tissue known as the indusium. Eastern Marsh Fern often occurs in saturated soils or even shallow water in full sunlight. It is sometimes available from nurseries. It requires a constantly moist soil and is one of the few ferns that thrive in direct sun. Eastern Marsh Fern is a vigorous grower wide spreading slender rhizomes. Under the right conditions, and can become “weedy” in the garden.--A. Diamond
Taxonomic Notes:**
Status:Native
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyTHELYPTERIDACEAE - Marsh Fern family
Genus Thelypteris
Species Thelypteris palustris Schott var. pubescens (G. Lawson) Fernald - Eastern Marsh Fern

Citation

Citation Thelypteris palustris Schott var. pubescens (G. Lawson) Fernald, Rhodora 31(362): 34. 1929.
Basionym: Lastrea thelypteris var. pubescens G. Lawson 1864.
Type: CANADA: Ontario: Addington Co.: Odessa, s.n., Lawson s.n. (holotype: ?).

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Dryopteris thelypteris subsp. pubescens Dryopteris thelypteris (Linnaeus) A. Gray, subsp. pubescens (G. Lawson) Hultén, Kongl. Svenska Vetenskapsakad. Handl. 8(5): 180. 1962.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Dryopteris thelypteris var. haleana Dryopteris thelypteris (Linnaeus) A. Gray, var. haleana (Fernald) M. Broun ex Weatherby, Amer. Fern J. 26: 95. 1936.Thelypteris palustris Schott, var. haleana Fernald 1929. 
Dryopteris thelypteris var. kamschatica Dryopteris thelypteris (Linnaeus) A. Gray, var. kamschatica C. Christensen, in Hultén, Fl. Kamtchat. 1: 38. 1927.  
Dryopteris tremula Dryopteris tremula Christ, Notul. Syst. (Paris) 1: 234. 1910. MEXICO: Michoacán: La Huerta, vicinity of Morelia, 15 Oct 1909, Arsène 3106 (holotype; P?; isotypes: F, GH, MEXU(3 shts), MO, NY, U).
Dryopteris thelypteris var. pubescens Dryopteris thelypteris (Linnaeus) A. Gray, var. pubescens (G. Lawson) Nakai, Bot. Mag. (Tokyo) 45: 97. 1931; (G. Lawson) Komarov, Izv. Bot. Sada Akad. Nauk SSSR 30: 193. 1932.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Filix-mas thelypteris var. pubescens Filix-mas thelypteris (Linnaeus) Farwell, var. pubescens (G. Lawson) Farwell, Amer. Midl. Naturalist 12: 258. 1931.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Lastrea thelypteris var. pubescens Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson, Edinburgh New Philos. J., ser. 2. 19: 277. 1864. CANADA: Ontario: Addington Co.: Odessa, s.n., Lawson s.n. (holotype: ?).
Thelypteris confluens var. pubescens Thelypteris confluens (Thunberg) C. V. Morton, var. pubescens (G. Lawson) Pringle, Techn. Bull. Roy. Bot. Gard. Hamilton 4: 42. 1969.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Thelypteris palustris subsp. pubescens Thelypteris palustris Schott, subsp. pubescens (G. Lawson) Holub, Folia Geobot. Phytotax. 4: 45. 1969.Lastrea thelypteris (Linnaeus) Bory, var. pubescens G. Lawson 1864. 
Thelypteris palustris var. haleana Thelypteris palustris Schott, var. haleana Fernald, Rhodora 31: 34, f. 5-6. 1929. LOUISIANA: Rapides Par.: Alexandria, s.d., Hale s.n. (holotype: GH).
Thelypteris thelypteris    

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Bibb, Butler, Choctaw, Clarke, Clay, Coffee, Conecuh, Covington, Crenshaw, Elmore, Escambia, Etowah, Lawrence, Lee, Limestone, Marengo, Mobile, Montgomery, Perry, Pike, Talladega, Tallapoosa, Tuscaloosa

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AMAL, AUA, JSU, TROY, UNA, USAM, UWAL

Range of years during which specimens were collected: -

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
UNA00057505 Clarke 25 Aug 1953 White, Jesse S.
s.n.
UNA
Thelypteris palustris Schott
AMAL00012385 Choctaw 03 Jul 1980 Kral, Robert
65913
AMAL
Thelypteris palustris Schott
TROY000013301 Baldwin 06 Jun 2006 Spaulding, Daniel D.
12467
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UWAL0047562 Baldwin 07 Aug 2015 Barger, T. Wayne
SP#1867
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
AUA000037671 Baldwin 4 Aug 1982 Short, John W.
1266
AUA
Thelypteris Palustris Schott
AUA_ACC_55130 Pike 7 Aug 1995 Diamond, Alvin R.
9824
AUA
Thelypteris Palustris Schott
UNA00052239 Talladega s.d. O'Kelley, Joseph C.
s.n.
UNA
Thelypteris palustris Schott
AUA_ACC_31772 Butler 15 Sep 1976 Short, John W.
819
AUA
Thelypteris Palustris Schott
UWAL0001419 Perry 07 Aug 2007 Keener, Brian R.
3836
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
AMAL00013561 Baldwin 06 Jun 2006 Spaulding, Daniel D.
12467
AMAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
AMAL00011934 Coffee 14 Nov 1978 Kral, Robert
63129
AMAL
Thelypteris palustris Schott
TROY000031556 Butler 15 Jul 2010 Diamond, Alvin R.
21462
TROY
Thelypteris palustris Schott
UNA00052252 Autauga 20 Jul 1976 Short, John W. (Jack)
772
UNA
Thelypteris palustris Schott
USAM000000641 Lawrence 4 Jul 2003 Pate, R.
72
USAM
Thelypteris palustris Schott
TROY000030804 Autauga 20 Jul 1976 Short, John W. (Jack)
772
TROY
Thelypteris palustris Schott
AMAL00017584 Escambia 17 Jun 2004 Diamond, Alvin R.
14442
AMAL
Thelypteris palustris Schott
UNA00063078 Covington 17 Jun 2004 Diamond, Alvin R.
14442
UNA
Thelypteris palustris Schott
UWAL0044027 Baldwin 25 Jun 2003 Larsen, Harry S.
1289
UWAL
Thelypteris palustris Schott var. pubescens (G. Lawson) Fernald
AUA_ACC_31773 Baldwin 2 Jun 1976 Short, John W.
601
AUA
Thelypteris Palustris Schott
UNA00068858 Perry 07 Aug 2007 Keener, Brian R.
3836
UNA
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
TROY000000478 Crenshaw 03 Jun 2001 Diamond, Alvin R.
12470
TROY
Thelypteris palustris Schott
UNA00052246 Mobile 22 May 1894 Mohr, Charles
s.n.
UNA
Thelypteris palustris Schott
TROY000014447 Perry 07 Aug 2007 Keener, Brian R.
3836
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UWAL0012711 Marengo 25 Jul 2011 England, J. Kevin
3097
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AUA_ACC_9959 Mobile 6 Jun 1961 Jones, Jr., Samuel B.
683
AUA
Thelypteris Palustris Schott
AMAL00020272 Baldwin 21 Apr 2011 Barger, T. Wayne
SP#311
AMAL
Thelypteris palustris Schott
UNA00052248 Baldwin 28 Apr 1947 Lueth, Francis X.
L 24
UNA
Thelypteris palustris Schott
USAM000000294 Baldwin 29 Jun 1974 LeLong, Michel G.
7992
USAM
Thelypteris palustris Schott
AUA_ACC_31769 Lee 21 Apr 1977 Short, John W.
890
AUA
Thelypteris Palustris Schott
AMAL00019903 Limestone 26 Jul 1983 Short, John W. (Jack)
1364
AMAL
Thelypteris palustris (A. Gray) Schott
UWAL0016925 Baldwin 13 Sep 2008 England, J. Kevin
1093
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000000475 Coffee 14 Nov 1978 Kral, Robert
63129
TROY
Thelypteris palustris Schott
ALNHS00003377 Baldwin 7 Aug 2015 Barger, T. Wayne
SP#1867
ALNHS
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UWAL0002857 Baldwin 09 Sep 2010 Diamond, Alvin R.
21560
UWAL
Thelypteris palustris Schott
AUA_ACC_8286 Baldwin 31 Aug 1960 Jones, Jr., Samuel B.
AUA
Thelypteris Palustris Schott
AMAL00009615 Baldwin 21 Apr 2011 Barger, T. Wayne
311
AMAL
Thelypteris palustris Schott
JSU109756 Clarke 19 Jul 1970 Kral, Robert
40191
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AUA_ACC_31842 Autauga 20 Jul 1976 Short, John W.
772
AUA
Thelypteris Palustris Schott
AMAL00019891 Baldwin 04 Aug 1982 Short, John W. (Jack)
1266
AMAL
Thelypteris palustris (A. Gray) Schott
TROY000040562 Marengo 25 Jul 2011 England, J. Kevin
3097
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UNA00052245 Autauga 28 May 1953 White, Jesse S.
s.n.
UNA
Thelypteris palustris Schott
JSU109759 Etowah 16 Sep 1995 Hodge-Spaulding, Lesley M.
4352
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000013477 Tallapoosa 11 Apr 2003 Spaulding, Daniel D.
11718
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
UNA00052247 Montgomery _ Oct 1879 Mohr, Charles
s.n.
UNA
Thelypteris palustris Schott
UWAL0001418 Baldwin 06 Jun 2006 Spaulding, Daniel D.
12467
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
AUA_ACC_71968 Baldwin 4 Aug 1982 Short, John W.
1262
AUA
Thelypteris Palustris Schott
ALNHS00000788 Baldwin 15 Sep 2011 Barger, T. Wayne
Perd-623
ALNHS
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
AUA_ACC_31841 Conecuh 16 Sep 1976 Short, John W.
826
AUA
Thelypteris Palustris Schott
UWAL0030070 Montgomery 10 Aug 2015 Diamond, Alvin R.
26745
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AUA_ACC_63823 Escambia 17 Jun 2004 Diamond, Alvin R.
14442
AUA
Thelypteris Palustris Schott
TROY000029603 Baldwin 16 Jun 2009 Barger, T. Wayne
623
TROY
Thelypteris palustris Schott
UWAL0017235 Lawrence 08 Jun 2014 England, J. Kevin
4918
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000000477 Crenshaw 27 Jul 2000 Diamond, Alvin R.
11928
TROY
Thelypteris palustris Schott
UNA00052253 Lee 21 Apr 1977 Short, John W. (Jack)
890
UNA
Thelypteris palustris Schott
AMAL00014169 Perry 07 Aug 2007 Keener, Brian R.
3836
AMAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
AUA_ACC_72231 Lawrence 8 Jun 2014 England, J. Kevin
4918
AUA
Thelypteris Palustris Schott Var. Pubescens (Lawson) Fernald
TROY000000479 Escambia 17 Jun 2004 Diamond, Alvin R.
14442
TROY
Thelypteris palustris Schott
JSU109757 Elmore 1 Aug 1978 Kral, Robert
62363
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AUA_ACC_31771 Covington 2 Jun 1976 Short, John W.
623
AUA
Thelypteris Palustris Schott
TROY000000474 Choctaw 03 Jul 1980 Kral, Robert
65913
TROY
Thelypteris palustris Schott
AMAL00019767 Baldwin 04 Aug 1982 Short, John W. (Jack)
1262
AMAL
Thelypteris palustris (A. Gray) Schott
TROY000000476 Conecuh 26 Aug 2000 Diamond, Alvin R.
12014
TROY
Thelypteris palustris Schott
ALNHS00000612 Baldwin 21 Apr 2011 Barger, T. Wayne
SP#311
ALNHS
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
JSU109758 Clay 9 Oct 1982 Whetstone, R. David
12715
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
USAM000000295 Mobile 6 Sep 1978 LeLong, Michel G.
10504
USAM
Thelypteris palustris Schott
TROY000043289 Lawrence 08 Jun 2014 England, J. Kevin
4918
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AMAL00005512 Pike 11 Jul 2001 Diamond, Alvin R.
12523
AMAL
Thelypteris palustris Schott
UWAL0030596 Covington 15 Aug 1987 Kral, Robert
74303
UWAL
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
UWAL0002113 Butler 15 Jul 2010 Diamond, Alvin R.
21462
UWAL
Thelypteris palustris Schott
TROY000042710 Montgomery 10 Aug 2015 Diamond, Alvin R.
26745
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AUA_ACC_31768 Mobile 2 Jun 1976 Short, John W.
590
AUA
Thelypteris Palustris Schott
UNA00052241 Tuscaloosa 11 May 1953 Chermock, Ralph L.
s.n.
UNA
Thelypteris palustris Schott
UNA00068909 Baldwin 06 Jun 2006 Spaulding, Daniel D.
12467
UNA
Thelypteris palustris Schott var. pubescens (Lawson) Fern.
TROY000033638 Barbour 01 Oct 2011 Morrison, Derek
129
TROY
Thelypteris palustris Schott
UNA00052244 Autauga 28 May 1953 White, Jesse S.
s.n.
UNA
Thelypteris palustris Schott
JSU109760 Etowah 16 Sep 1995 Hodge-Spaulding, Lesley M.
4352
JSU
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
TROY000043989 Bibb 07 Aug 2015 England, J. Kevin
5570
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
AMAL00005511 Mobile 05 Jun 1953 Brown, J.S.
s.n.
AMAL
Thelypteris palustris Schott
UWAL0030421 Bibb 07 Aug 2015 England, J. Kevin
5570
UWAL
Thelypteris palustris (Schott) var. pubescens (Lawson) Fernald
AUA_ACC_34605 Clarke 8 May 1976 Short, John W.
506
AUA
Thelypteris Palustris Schott
TROY000032980 Baldwin 09 Sep 2010 Diamond, Alvin R.
21560
TROY
Thelypteris palustris Schott
AUA_ACC_31770 Escambia 16 Sep 1976 Short, John W.
831
AUA
Thelypteris Palustris Schott
UNA00052240 Choctaw 05 Aug 1953 Chermock, Ralph L.
s.n.
UNA
Thelypteris palustris Schott
UNA00056167 Mobile 05 Jun 1953 Brown, Jack S.
s.n.
UNA
Thelypteris palustris Schott
TROY000000480 Pike 11 Jul 2001 Diamond, Alvin R.
12523
TROY
Thelypteris palustris Schott var. pubescens (Lawson) Fernald
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