Helenium amarum

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Distribution Map: Based on vouchered plant specimens only. View county names by placing the mouse cursor over a particular county.

Species Distribution Map
Family:ASTERACEAE
Species:Helenium amarum (Raf.) H. Rock
Common Name:Bitterweed; Yellowdicks; Five Leaf Sneezeweed; Spanish Daisy
Habitat:Disturbed soils.
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Annual
Category:Vascular
USDA Symbol:HEAM
Plant Notes:Bitterweed is a native herbaceous annual species in the Sunflower family (Asteraceae). It occurs throughout Alabama. Bitterweed grows in disturbed areas such as on roadsides, in urban areas, and in fields and pastures. It is an annual with a tap root. Stems are erect, usually branched, green in color, and glabrous or slightly pubescent. Leaves are alternate, sessile, linear in outline, entire or with a few small teeth, and glabrous or with a few hairs. The lower stem leaves are often withered at flowering. All parts of the plants release an unpleasant odor when damaged. Flowers are produced in heads. The heads are arranged in paniculiform arrays. Each head has 8-10 yellow ray flowers and 75-250 yellow disc flowers. The fruit is an achene. The common name is in reference to this plant causing milk to taste bitter when cows graze on the foliage. A large population of Bitterweed in a pasture is often a sign of overgrazing. Bitterweed is an easy to grow annual that is tolerant of poor soils and dry conditions. It can be used for borders, rock gardens, and for xeriscape gardening. It has an extended blooming period and is an important pollinator plant, attracting butterflies, sweat bees, native wasps, and beetles. All parts of the plant contain the glycoside gudaldin and can cause toxic symptoms. A decoction of the entire plan was used to make “Bitterweed Tea”, a folk remedy for intestinal worms.—A. Diamond.
Taxonomic Notes:**
Status:Native, FACU- (NWI)
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyASTERACEAE - Aster family
Genus Helenium
Species Helenium amarum (Raf.) H. Rock - Bitterweed; Yellowdicks; Five Leaf Sneezeweed; Spanish Daisy

Citation

Citation Helenium amarum (Rafinesque) H. Rock, Rhodora 59(702): 131. 1957.
Basionym: Gaillardia amara Raf. 1817.
Type: USA: LOUISIANA: Rapides Par.: Alexandria, 6 Sep 1898, Ball 182 (neotype: US). Neotypified by H. Rock, Rhodora 59: 131. 1957.

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Gaillardia amara Gaillardia amara Rafinesque, Fl. Ludov. 69. 1817. LOUISIANA: Rapides Par.: Alexandria, 6 Sep 1898, Ball 182 (neotype: US). Neotypified by H. Rock, Rhodora 59: 131. 1957.
Heleniastrum tenuifolium Heleniastrum tenuifolium (Nuttall) Kuntze, Revis. Gen. Pl. 1: 342. 1891.Helenium tenuifolium Nuttall 1834. 
Helenium tenuifolium Helenium tenuifolium Nuttall, J. Acad. Nat. Sci. Philadelphia 7: 66. 1834. ALABAMA: Without data, Beck s.n. (lectotype: PH). Lectotypified by Bierner, Sida 13: 457. 1989.

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Bibb, Blount, Bullock, Butler, Calhoun, Chambers, Cherokee, Chilton, Choctaw, Clarke, Clay, Cleburne, Coffee, Colbert, Conecuh, Coosa, Covington, Crenshaw, Cullman, Dale, Dallas, DeKalb, Elmore, Escambia, Etowah, Fayette, Franklin, Geneva, Greene, Hale, Henry, Houston, Jackson, Jefferson, Lamar, Lauderdale, Lawrence, Lee, Limestone, Lowndes, Macon, Madison, Marengo, Marion, Marshall, Mobile, Monroe, Montgomery, Morgan, Perry, Pickens, Pike, Randolph, Russell, Shelby, St. Clair, Sumter, Talladega, Tallapoosa, Tuscaloosa, Walker, Wilcox, Winston

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AMAL, AUA, JSU, SAMF, TROY, UNA, USAM, UWAL

Range of years during which specimens were collected: 1949 - 2023

Barcode / Accession No. County Coll. Date Collector &
Collection No.		 Herbarium &
Herbarium Name Used		 Image
JSU111766 Cherokee 10 Jul 1988 Dickson, Catherine C.
1903		 JSU
Helenium amarum (Rafinesque) H. Rock
AMAL00003023 Walker 20 Jul 2002 Spaulding, Daniel D.
11451		 AMAL
Helenium amarum (Raf.) H. Rock var. amarum
AUA000004222 Coosa 10 Sep 1975 Rutland, David W.
1269		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004224 Coosa 2 Sep 1975 Rutland, David W.
1080		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0036740 Winston 05 Aug 2017 England, J. Kevin
8334		 UWAL
Helenium amarum (Rafinesque) H. Rock var. amarum
TROY000015700 Lamar 28 Jul 2002 Keener, Brian R.
2319		 TROY
Helenium amarum (Rafinesque) H. Rock
AUA000004279 Russell 18 Jun 2002 Gil, Glenda
2001-246		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004277 Randolph 24 Jun 1986 Nixon, C.F.
2249		 AUA
Helenium Amarum (Raf.) H. Rock
SAMF0005453 Bibb 29 Jul 2011 Davenport, Lawrence J.
4738		 SAMF
Helenium amarum (Rafinesque) H. Rock
JSU111768 Cleburne 30 May 1972 Whetstone, R. David
RDW 77		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004245 Henry 7 Jul 1988 Diamond, Alvin R.
5158		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000683 Sumter 01 Jul 2000 Holland, Richard
AP00-52		 UWAL
Helenium amarum (Raf.) H. Rock
UNA00046879 Greene 15 Aug 1965 Naugle, Helen E.
G64		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002163 Crenshaw 09 Jul 2000 Diamond, Alvin R.
11787		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111763 Calhoun 7 Jul 1968 Johnson, Stephen C.
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004198 Chilton 21 Aug 1975 Rutland, David W.
820		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004208 Clay 24 Jun 1975 Rutland, David W.
651		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004220 Coosa 3 Sep 1975 Rutland, David W.
1182		 AUA
Helenium Amarum (Raf.) H. Rock
AMAL00003016 Autauga 07 Sep 2002 Spaulding, Daniel D.
11566		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004291 Tuscaloosa 9 Jul 1970 Moore, Harold D.
283		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000015703 Randolph 06 Oct 1983 Nixon, Christopher F.
489		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111781 Etowah 30 Aug 1968 Clonts, Jerry A.
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111805 Winston 7 Jun 1975 Whetstone, R. David
4526		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004228 Crenshaw 4 Jun 1995 Diamond, Alvin R.
9575		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004213 Cleburne 12 Sep 1975 Rutland, David W.
1798		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000684 Talladega 19 Oct 2007 Keener, Brian R.
3968		 UWAL
Helenium amarum (Raf.) H. Rock
TROY000041982 Macon 07 Nov 2014 Diamond, Alvin R.
25659		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111796 Marshall 2 Aug 1974 Atkinson, Tim A.
393		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004261 Lee 1 Jul 1966 Daly, Jr., Robert
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004292 Wilcox 29 Aug 1960 Jones, Jr., Samuel B.
215		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046888 Lee 07 Sep 1980 Latham, B.
17		 UNA
Helenium amarum (Raf.) H. Rock
TROY000038483 Dallas 01 Sep 2012 Boozer, James
5		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111801 Talladega 13 Jul 1992 Ballard, J. Mark
335		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004233 Dallas 25 Oct 1987 Diamond, Alvin R.
3741		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004206 Clay 24 Jun 1975 Rutland, David W.
627		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004237 Elmore 28 Jul 1975 Rutland, David W.
737		 AUA
Helenium Amarum (Raf.) H. Rock
USAM000002314 Mobile 04 Aug 1967 LeLong, Michel G.
3483		 USAM
Helenium amarum (Rafinesque) H. Rock
JSU111794 Marion 4 Oct 1980 Whetstone, R. David
10377		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004252 Lee Belcher, C.C.
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004215 Cleburne 11 Sep 1975 Rutland, David W.
1643		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004200 Chilton 21 Aug 1975 Rutland, David W.
806		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046882 Tuscaloosa 26 Jul 1957 Coffman, L.F.
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
AUA000004234 Dallas 7 Sep 1970 Moore, Harold D.
849		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00059509 Autauga 07 Sep 2002 Spaulding, Daniel D.
11566		 UNA
Helenium amarum (Raf.) H. Rock
JSU111791 Lawrence 12 Aug 1974 Atkinson, Tim A.
664		 JSU
Helenium amarum (Rafinesque) H. Rock
AMAL00017680 Lauderdale 25 Jun 2009 Holt, Brian D.
SC-99		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004271 Monroe 18 Jun 1972 Hybart, Michael G.
018		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0051308 Sumter 17 May 1969 Dean, Blanche E.
247		 UWAL
Helenium amarum
TROY000015702 Tallapoosa 10 Aug 2006 Barger, T. Wayne
314		 TROY
Helenium amarum (Rafinesque) H. Rock
TROY000015805 Marengo 24 Sep 2008 England, J. Kevin
1263		 TROY
Helenium amarum (Rafinesque) H. Rock var. amarum
JSU111786 Jefferson 17 Sep 1983 Whetstone, R. David
12452		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004196 Chilton 21 Aug 1975 Rutland, David W.
865		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004285 Talladega 28 Aug 1975 Rutland, David W.
984		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0019952 Morgan 15 Aug 2015 England, J. Kevin
5602		 UWAL
Helenium amarum (Rafinesque) H.Rock
USAM000002313 Greene 07 Jun 1979 LeLong, Michel G.
10828		 USAM
Helenium amarum (Rafinesque) H. Rock
JSU111773 Cullman 5 Aug 1974 Whetstone, R. David
3411		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004265 Macon 6 Oct 1971 Boone, Allison
117		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004214 Cleburne 12 Sep 1975 Rutland, David W.
1736		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000038890 Cullman Wolf, Wolfgang
1339		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046891 Mobile 08 Apr 1950 Crawford, Lloyd C.
853		 UNA
Helenium amarum (Raf.) H. Rock
AUA000004202 Choctaw 8 Sep 1970 Moore, Harold D.
976		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000002171 Pike 18 Oct 1995 Holmes, Jeanese
78		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111804 Walker 13 Aug 1974 Atkinson, Tim A.
685		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004248 Jefferson 29 Jul 1967 Carter, Richard D.
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004246 Henry 13 Aug 1960 Jones, Jr., Samuel B.
245		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0037320 Bullock 18 Jun 2016 Soehren, Eric C.
344		 UWAL
Helenium amarum (Raf.) H.Rock var. amarum
TROY000039818 Baldwin 13 Mar 2013 Barger, T. Wayne
1148		 TROY
Helenium amarum (Rafinesque) H. Rock
TROY000021582 Walker 20 Jul 2002 Keener, Brian R.
2278		 TROY
Helenium amarum (Rafinesque) H. Rock
AMAL00003017 Blount 07 Jul 1997 Keener, Brian R.
220		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004207 Clay 23 Jun 1975 Rutland, David W.
504		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004236 Elmore 28 Jul 1975 Rutland, David W.
668		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0017293 Lawrence 15 Sep 2013 England, J. Kevin
4539		 UWAL
Helenium amarum (Rafinesque) H. Rock
JSU111779 Etowah 30 Aug 1968 Clonts, Jerry A.
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111760 Blount 7 Aug 1975 Whetstone, R. David
5931		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004225 Covington 3 Oct 1987 Diamond, Alvin R.
3681		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004212 Cleburne 12 Sep 1975 Rutland, David W.
1765		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000685 Walker 20 Jul 2002 Keener, Brian R.
2278		 UWAL
Helenium amarum (Raf.) H. Rock
UNA00053862 Blount 07 Jul 1997 Keener, Brian R.
220		 UNA
Helenium amarum (Raf.) H. Rock
JSU111783 Jackson 27 Jun 1998 Brodeur, Kristin R.
841		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004243 Hale 8 Sep 1970 Moore, Harold D.
983		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000002164 Dale 15 May 1995 Jernigan, Clay
7		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111798 Morgan 14 Aug 1974 Atkinson, Tim A.
775		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004203 Choctaw 26 Jul 1994 Crouch, Virginia E.
1106		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004255 Lee 26 Sep 1959 Jones, Jr., Samuel B.
 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0052921 Bibb 28 Jun 2019 Keener, Brian R.
11412		 UWAL
Helenium amarum (Raf.) H. Rock
TROY000030364 Jackson 27 Aug 2009 Barger, T. Wayne
380		 TROY
Helenium amarum (Rafinesque) H. Rock
TROY000033461 Barbour 03 Sep 2011 Ryan, Lauren
80		 TROY
Helenium amarum (Raf.) H. Rock
AMAL00003019 Lamar 28 Jul 2002 Keener, Brian R.
2319		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004210 Clay 24 Jun 1975 Rutland, David W.
591		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004235 Elmore 28 Jul 1975 Rutland, David W.
682		 AUA
Helenium Amarum (Raf.) H. Rock
JSU143476 Etowah 28 Jun 1969 Cofield, Jan
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111800 St. Clair 9 Sep 1983 Whetstone, R. David
13722		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004221 Coosa 10 Sep 1975 Rutland, David W.
1248		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004276 Randolph 16 Jun 1975 Rutland, David W.
325		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0052066 Madison 04 Oct 2020 England, J. Kevin
11329		 UWAL
Helenium amarum (Raf.) H. Rock
TROY000002170 Monroe 26 Aug 2006 Diamond, Alvin R.
16822		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111780 Etowah 19 Jun 1994 Hodge, Lesley M.
2661		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004273 Pike 7 Aug 1987 Diamond, Alvin R.
3463		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004268 Macon 12 Jul 1969 Simonds, E.P.
007		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046889 Lee 18 Jun 1972 Moody, Gary
18		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002161 Covington 08 Jul 2001 Diamond, Alvin R.
12497		 TROY
Helenium amarum (Rafinesque) H. Rock
AUA000004260 Lee 11 Aug 1983 Fleischer, S.J.
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004253 Lee 28 May 1970 Murchison, Grover R.
201		 AUA
Helenium Amarum (Raf.) H. Rock
JSU111770 Clay 19 Oct 1996 Ballard, J. Mark
7013		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000025172 Covington 20 Jun 2008 Hansen, Curtis J.
3375		 AUA
Helenium Amarum (Raf.) H. Rock Var. Amarum
AUA000004209 Clay 24 Jun 1975 Rutland, David W.
639		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004192 Calhoun 29 Aug 1975 Rutland, David W.
989		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0056915 Bibb 05 Jun 2019 Keener, Brian R.
11301		 UWAL
Helenium amarum (Rafinesque) H.Rock
JSU111795 Marshall 9 Sep 1991 Spaulding, Daniel D.
831		 JSU
Helenium amarum (Rafinesque) H. Rock
AMAL00003018 Calhoun 20 Jun 1989 Spaulding, Daniel D.
225		 AMAL
Helenium amarum (Raf.)
AUA000004223 Coosa 10 Sep 1975 Rutland, David W.
1289		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004219 Coosa 2 Sep 1975 Rutland, David W.
1149		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0046233 Autauga 28 Jun 2017 Parker, Matthew R.
970		 UWAL
Helenium amarum (Rafinesque) H. Rock
TROY000002167 Dale 08 May 1997 Rundell, Hannelore
145		 TROY
Helenium amarum (Rafinesque) H. Rock
AUA000004275 Pike 31 Jul 1970 Moore, Harold D.
554		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000679 Covington 02 Aug 2008 Diamond, Alvin R.
19679		 UWAL
Helenium amarum (Raf.) H. Rock
JSU111778 Etowah 10 Aug 1974 Atkinson, Tim A.
608		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111802 Talladega 30 May 1973 Whetstone, R. David
RDW-938		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004238 Escambia 7 Nov 1987 Diamond, Alvin R.
3764		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0051309 Jefferson 07 Sep 1984 Barber, J. Patrick
1087		 UWAL
Helenium amarum (Raf.) H. Rock
UNA00046880 Greene 17 Sep 1966 Naugle, Helen E.
G176		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002246 Covington 17 Sep 2005 Diamond, Alvin R.
15780		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111765 Calhoun 15 Jun 1982 Whetstone, R. David
11307		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004199 Chilton 21 Aug 1975 Rutland, David W.
839		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004283 Talladega 28 Aug 1975 Rutland, David W.
926		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0007857 Pike 26 Sep 2010 Dykes, Bailee
1		 UWAL
Helenium amarum (Raf.) H. Rock
JSU111761 Calhoun 20 Jun 1969 Tudor, Jessica
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004266 Macon 13 May 1967 Barrick, William E.
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004211 Cleburne 11 Sep 1975 Rutland, David W.
1457		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004197 Chilton 21 Aug 1975 Rutland, David W.
792		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046886 Hale 07 Sep 1952 Chermock, Ralph L.
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002166 Dale 18 Jul 1997 Rundell, Hannelore
293		 TROY
Helenium amarum (Rafinesque) H. Rock
AUA000004201 Chilton 7 Jul 1970 Moore, Harold D.
200		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00058652 Lamar 28 Jul 2002 Keener, Brian R.
2319		 UNA
Helenium amarum (Raf.) H. Rock
JSU111772 Cullman 12 Aug 1974 Atkinson, Tim A.
621		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111759 Bibb 31 Jul 1998 Whetstone, R. David
19598		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004287 Tallapoosa 24 Jun 1974 Hutto, T.A.
378		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0050771 Jefferson 23 Sep 1984 Bussey, Melanie G.
980		 UWAL
Helenium amarum (Raf.) H. Rock
UNA00046892 Randolph 20 Jul 1986 Nixon, Christopher F.
2406		 UNA
Helenium amarum (Raf.) H. Rock
TROY000015227 Geneva 28 Aug 2008 Diamond, Alvin R.
19853		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111806 Winston 17 Aug 1974 Whetstone, R. David
3836		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004195 Chilton 21 Aug 1975 Rutland, David W.
860		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004284 Talladega 28 Aug 1975 Rutland, David W.
948		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0010422 Bullock 18 Sep 2011 Dykes, Bailee
132		 UWAL
Helenium amarum (Raf.) H. Rock
UWAL0056172 Fayette 19 Jun 2023 England, Kevin J.
13754		 UWAL
Helenium amarum (Rafinesque) H. Rock
AUA000004274 Pike 15 Jul 1988 Diamond, Alvin R.
5214		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000681 Marengo 24 Sep 2008 England, J. Kevin
1263		 UWAL
Helenium amarum (Raf.) H. Rock
UNA00066283 Monroe 26 Aug 2006 Diamond, Alvin R.
16822		 UNA
Helenium amarum (Raf.) H. Rock
JSU111790 Jefferson 28 May 1984 Barber, J. Patrick
419		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111774 DeKalb 11 Jul 1993 Spaulding, Daniel D.
5011		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004264 Lowndes 6 Jul 1967 Atkins, W.B.
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004188 Autauga 15 Jul 1970 Moore, Harold D.
382		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000682 Perry 07 Nov 2008 England, J. Kevin
1488		 UWAL
Helenium amarum (Raf.) H. Rock
TROY000014533 Barbour 17 Jul 2008 Dyess, Kaleb K.
1		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111784 Jackson 21 Sep 1997 Spaulding, Daniel D.
9984		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004229 Dale 23 Jun 1991 Diamond, Alvin R.
7518		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00060553 Mobile 01 Jun 1965 Deramus, Rebecca
497		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002159 Coffee 14 Oct 2000 Windham, Mark
3		 TROY
Helenium amarum (Rafinesque) H. Rock
AMAL00015044 Houston 19 Nov 2008 Barger, T. Wayne
SP#44		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004205 Clarke 4 Sep 1971 Henley, Nancy S.
013		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00073038 Houston 19 Jul 2016 Barger, T. Wayne
DTP-291		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002168 Houston 02 Sep 2004 Diamond, Alvin R.
14658		 TROY
Helenium amarum (Rafinesque) H. Rock
USAM000002316 Baldwin 10 Aug 1980 LeLong, Michel G.
11309		 USAM
Helenium amarum (Rafinesque) H. Rock
AUA000004189 Barbour 17 Jul 1988 Diamond, Alvin R.
5224		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0036484 Greene 28 Oct 2017 England, J. Kevin
8615		 UWAL
Helenium amarum (Rafinesque) H. Rock var. amarum
UNA00069576 Henry 24 Jul 2008 Dyess, Kaleb K.
26		 UNA
Helenium amarum (Raf.) H. Rock
JSU111785 Jefferson 7 Sep 1984 Barber, J. Patrick
1258		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU139723 Limestone 13 Jul 1996 Spaulding, Daniel D.
9072		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004250 Lee 10 Oct 1999 Hansen, Curtis J.
99-29		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004242 Geneva 18 Jul 1970 Moore, Harold D.
637		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046885 St. Clair 24 Jun 1951 Hollis, C.G.
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
TROY000020021 Dale 31 Jul 2008 Dyess, Kaleb K.
57		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111792 Limestone 27 Jun 1996 Whetstone, R. David
17905		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004270 Mobile 15 Aug 1973 Kral, Robert
51351		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046878 Marengo 07 Oct 1967 Thomas, Joab L.
1241		 UNA
Helenium amarum (Raf.) H. Rock
AUA000004218 Conecuh 12 Jul 1985 Diamond, Alvin R.
1173		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000039387 Tallapoosa 28 Oct 1966 Hocking, George M.
743		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00060905 Mobile 25 Dec 2002 Zebryk, Tad M.
8181		 UNA
Helenium amarum (Raf.) H. Rock
TROY000048712 Autauga 28 Jun 2017 Parker, Matthew R.
970		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111769 Clarke 13 Sep 1997 Whetstone, R. David
18750		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004240 Franklin 13 Jun 1972 Pounders, C.T.
043		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0036721 Lamar 13 Jul 2017 England, J. Kevin
8113		 UWAL
Helenium amarum (Rafinesque) H. Rock var. amarum
TROY000002162 Crenshaw 01 Nov 2003 Cho, I.K.
2		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111776 DeKalb 1 Sep 1982 Black, Randall
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
AMAL00017123 Shelby 29 Sep 1999 Morrow. D. Keener
875		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004263 Lowndes 10 Jul 1970 Moore, Harold D.
1137		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004241 Geneva 25 Jul 1969 Moore, Harold D.
437-69		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000015224 Monroe 20 Sep 2007 Diamond, Alvin R.
18241		 TROY
Helenium amarum (Rafinesque) H. Rock
SAMF0005454 Perry 03 Nov 2005 Davenport, Lawrence J.
3384		 SAMF
Helenium amarum (Rafinesque) H. Rock
AUA000004194 Chambers 23 Aug 1991 Diamond, Alvin R.
7698		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004286 Tallapoosa 20 Jun 1969 Thornhill, Randy
002		 AUA
Helenium Amarum (Raf.) H. Rock
ALNHS00002332 Colbert 22 Aug 2013 Barger, T. Wayne
FH-104		 ALNHS
Helenium amarum (Raf.) H. Rock var. amarum
JSU111799 Randolph 5 Jul 1969 Herren, Tommy
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000037556 Morgan 15 Aug 2015 England, J. Kevin
5602		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004251 Lee 1 Jul 1948 Searcy, V.S.
 AUA
Helenium Amarum (Raf.) H. Rock
TROY000046450 Lowndes 19 Jun 2018 Diamond, Alvin R.
28374		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111797 Morgan 14 Aug 1974 Whetstone, R. David
3718		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004256 Lee 15 Jul 1969 Rebois, R.V.
038		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00048690 Houston 02 Sep 1988 Burckhalter, Robert
1489		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002160 Coffee 19 Jun 1999 Martin, Brian H.
49		 TROY
Helenium amarum (Rafinesque) H. Rock
SAMF0005456 Mobile 23 Sep 1989 Green, LeAnne
35		 SAMF
Helenium amarum (Rafinesque) H. Rock
JSU111764 Calhoun 23 Jun 1985 Boozer, B.
005		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004258 Lee 11 Jul 1957 Arner, Dale H.
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004272 Monroe 10 Sep 1970 Moore, Harold D.
1081		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004226 Covington 25 Jul 1969 Moore, Harold D.
454-69		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000002172 Wilcox 21 Aug 2005 Diamond, Alvin R.
15665		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111787 Jefferson 23 Sep 1984 Bussey, Melanie G.
980		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004216 Coffee 23 Jul 1989 Diamond, Alvin R.
6053		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004278 Russell 21 Jun 1969 Weaver, Jimmy
021		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000002169 Montgomery 15 Oct 2000 Diamond, Alvin R.
12207		 TROY
Helenium amarum (Rafinesque) H. Rock
AMAL00013039 Limestone 19 May 1989 Alvis, Amy
008		 AMAL
Helenium amarum (Raf.) H. Rock var. amarum
AUA000004254 Lee 10 Jul 1948 Heinselman
 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004249 Jefferson 7 Jul 1977 Sessler, Ann H.
1688		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000033808 Bullock 18 Sep 2011 Dykes, Bailee
132		 TROY
Helenium amarum (Rafinesque) H. Rock
SAMF0005452 Shelby 12 Oct 2010 Davenport, Lawrence J.
4556		 SAMF
Helenium amarum (Rafinesque) H. Rock
AUA000004232 Dale 16 Aug 1991 Diamond, Alvin R.
7679		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0030245 Macon 07 Nov 2014 Diamond, Alvin R.
25659		 UWAL
Helenium amarum (Rafinesque) H.Rock
UNA00069037 Barbour 17 Jul 2008 Dyess, Kaleb K.
1		 UNA
Helenium amarum (Raf.) H. Rock
JSU111788 Jefferson 7 Sep 1984 Barber, J. Patrick
1087		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111793 Madison 11 May 1996 Spaulding, Daniel D.
8294		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004288 Tallapoosa 23 Sep 1972 Hagan, Patrick D.
032		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004282 Sumter 16 Jun 1970 Moore, Harold D.
077		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0043163 Baldwin 22 Jul 2003 Larsen, Harry S.
1412		 UWAL
Helenium amarum (Raf.) H. Rock
TROY000028048 Henry 24 Jul 2008 Dyess, Kaleb K.
26		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111803 Talladega 25 May 1972 Whetstone, R. David
RDW 43		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004187 Autauga 13 Aug 1982 Gunn, Scott C.
1008		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046877 Hale 14 Jun 1965 Maginness, Sherrie E.
13		 UNA
Helenium amarum (Raf.) H. Rock
AMAL00003021 Madison 14 Jul 1996 Threlkeld, Steven J.
724		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004191 Butler 19 Jun 1971 Hester, D.
015		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046883 Hale 25 Sep 1965 Williams, Norris H.
142		 UNA
Helenium amarum (Raf.) H. Rock
TROY000015223 Covington 22 Dec 2007 Diamond, Alvin R.
18691		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111789 Jefferson 13 Jul 1984 Barber, J. Patrick
761		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004227 Covington 17 Sep 2005 Diamond, Alvin R.
15780		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0036467 Tuscaloosa 30 Sep 2017 England, J. Kevin
8536		 UWAL
Helenium amarum (Rafinesque) H. Rock var. amarum
UNA00046887 Tuscaloosa 08 Jun 1950 Williams, B.C.
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
JSU111777 DeKalb 5 Jul 1969 Beaty, Norma J.
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111767 Chilton 22 Jul 1982 Whetstone, R. David
11733		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004193 Chambers 11 Jul 2000 Hansen, Curtis J.
2000-26		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004269 Marengo 19 Jul 1969 Moore, Harold D.
274-69		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00026665 Pickens 19 Sep 1985 Horn, Charles N.
981		 UNA
Helenium amarum (Raf.) H. Rock
TROY000002165 Dale 09 Aug 1999 Pennington, Tiffany
39		 TROY
Helenium amarum (Rafinesque) H. Rock
AMAL00006306 Monroe 26 Aug 2006 Diamond, Alvin R.
16822		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004217 Colbert 17 Jul 1965 Landers, Ken
 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046881 Tuscaloosa 07 Sep 1966 Thomas, Joab L.
864		 UNA
Helenium amarum (Raf.) H. Rock
USAM000002312 Mobile 14 Jul 2003 Pate, Ab R.
58		 USAM
Helenium amarum (Rafinesque) H. Rock
AUA000004290 Tallapoosa 25 Oct 1975 Dupree, W.G.
130		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004267 Macon 10 Jul 1970 Hopson, George R.
.0062		 AUA
Helenium Amarum (Raf.) H. Rock
ALNHS00000383 Dallas 24 May 2010 Barger, T. Wayne
OC-246		 ALNHS
Helenium amarum (Raf.) H. Rock var. amarum
UNA00026681 Mobile _ Sep 1871 Mohr, Charles
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
AMAL00003022 Montgomery 15 Oct 2000 Diamond, Alvin R.
12207		 AMAL
Helenium amarum (Raf.) Rock
AUA000004289 Tallapoosa 30 Jul 1972 Roboski, J.C.
121		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00046890 Tuscaloosa 06 Aug 1949 Carlin, May Gill
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
SAMF0006836 Mobile 13 Apr 2019 Hatcher, Joy
10		 SAMF
Helenium amarum (Linnaeus) H. Rock
JSU111762 Calhoun 23 Jun 1985 Boozer, B.
005		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004247 Houston 9 Aug 1969 Atkeson, Thomas D.
279		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004257 Lee 12 May 1969 Moore, Harold D.
216-89		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004239 Escambia 26 Aug 1969 Moore, Harold D.
618-69		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000015226 Perry 21 Jul 2003 Diamond, Alvin R.
14036		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU146188 Calhoun 10 Oct 1997 Hutchinson, Francine
2506		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004230 Dale 23 Jul 1989 Diamond, Alvin R.
6074		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004262 Limestone 3 Jul 1971 White, Dudley
008		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000002158 Coffee 29 Sep 2004 Diamond, Alvin R.
14730		 TROY
Helenium amarum (Rafinesque) H. Rock
AMAL00019414 Calhoun 25 Jul 1994 Ballard, J. Mark
4155		 AMAL
Helenium amarum (Raf.) H. Rock
AUA000004281 Sumter 29 Jun 1981 Henderson, T.
244		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0000680 Covington 22 Dec 2007 Diamond, Alvin R.
18691		 UWAL
Helenium amarum (Raf.) H. Rock
TROY000015701 Autauga 07 Sep 2002 Spaulding, Daniel D.
11566		 TROY
Helenium amarum (Rafinesque) H. Rock
TROY000048117 Autauga 28 Jul 2017 Parker, Matthew R.
1068		 TROY
Helenium amarum (Rafinesque) H. Rock
JSU111771 Colbert 17 Jul 1965 Landers, Kenneth E.
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004231 Dale 19 Jul 1989 Diamond, Alvin R.
6000		 AUA
Helenium Amarum (Raf.) H. Rock
UWAL0039698 Lowndes 19 Jun 2018 Diamond, Alvin R.
28374		 UWAL
Helenium amarum (Raf.) H. Rock
UNA00046884 Hale 08 Jun 1966 Williams, Norris H.
274		 UNA
Helenium amarum (Raf.) H. Rock
JSU111782 Jackson 5 Jul 1969 Bible, Rebecca
s.n.		 JSU
Helenium amarum (Rafinesque) H. Rock
JSU111775 DeKalb 23 Jul 1975 Whetstone, R. David
5257		 JSU
Helenium amarum (Rafinesque) H. Rock
AUA000004204 Clarke Maddox, William K.
009		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004244 Henry 19 Jul 1970 Moore, Harold D.
456		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00057163 Jackson 21 Sep 1997 Spaulding, Daniel D.
9984		 UNA
Helenium amarum (Raf.) H. Rock
SAMF0005455 Morgan 02 Aug 2005 Davenport, Lawrence J.
3304		 SAMF
Helenium amarum (Rafinesque) H. Rock
AUA000004190 Butler 2 Aug 1985 Diamond, Alvin R.
1324		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000004259 Lee 13 Sep 1971 Little, Jackie A.
026		 AUA
Helenium Amarum (Raf.) H. Rock
UNA00026682 s.d. Unknown
s.n.		 UNA
Helenium amarum (Raf.) H. Rock
USAM000002315 Madison 28 Jul 1976 Short, Nancy
s.n.		 USAM
Helenium amarum (Rafinesque) H. Rock
AUA000004280 St. Clair 30 Sep 1972 Eads, Beth
021		 AUA
Helenium Amarum (Raf.) H. Rock
AUA000028974 Clarke 22 May 2012 Horne, Howard E.
SWS-180		 AUA
Helenium Amarum (Raf.) H. Rock
TROY000015225 Escambia 18 Sep 2008 Diamond, Alvin R.
20060		 TROY
Helenium amarum (Rafinesque) H. Rock
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