Echinochloa colona

Family:POACEAE
Species:Echinochloa colona (L.) Link
Common Name:Jungle Rice; Shama Millet; Awnless Barnyard Grass
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Annual
Category:Vascular
USDA Symbol:**
Plant Notes:**
Taxonomic Notes:Ward (2005) made an argument that the spelling of the epithet should be "colonum" like the basionym. APA will follow IPNI and at this time IPNI uses "colona".
Status:Not Native, FACW (NWI)
References:Ward, D.B. 2005. A case of disputed orthography: is it Echinochloa colona; or is it Echinochloa colonum (Gramineae). Sida 21: 2171-2183.
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyPOACEAE - Grass family
Genus Echinochloa
Species Echinochloa colona (L.) Link - Jungle Rice; Shama Millet; Awnless Barnyard Grass

Citation

Citation Echinochloa colona (Linnaeus) Link, Hort. Berol. [Link] 2: 209. 1833.
Basionym: Panicum colonum L. 1759.
Type: JAMAICA: Without data, Browne s.n. (lectotype: LINN 80.23). Lectotypified by Clayton & Renvoize, in Polhill, Fl. Trop. E. Afr., Gramin. 557. 1982.

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Brachiaria longifolia Brachiaria longifolia Gilli, Ann. Naturhist. Mus. Wien 69: 39. 1966. TANZANIA: Njombe Distr.: Lupingu, s.d., Gilli 43 (holotype: W).
Echinochloa colona var. equitans Echinochloa colona (Linnaeus) Link, var. equitans (A. Richard) Cufodontis, Bull. Jard. Bot. Etat 39(Suppl.): 1302. 1969.Panicum equitans A. Richard 1850 
Echinochloa colona var. zonalis Echinochloa colona (Linnaeus) Link, var. zonalis (Gussone) Wooton & Standley, New Mexico Agric. Exp. Sta. Press Bull. 81: 45. 1912.Panicum zonale Gussone 1827. 
Echinochloa crus-galli subsp. colona Echinochloa crus-galli (Linnaeus) Palisot de Beauvois, subsp. colona (Linnaeus) Honda, Bot. Mag. (Tokyo) 37: 122. 1923.Panicum colonum Linnaeus 1759. 
Echinochloa crusgalli var. longiseta Echinochloa crusgalli (Linnaeus) Palisot de Beauvois, var. longiseta (Trinius) Hara, Bot. Mag. (Tokyo) 52: 231. 1938, non (Döll) Podpera 1926.Panicum crusgalli Linnaeus, var. longisetum Trinius 1829. 
Echinochloa divaricata Echinochloa divaricata Andersson, in Peters, Naturwiss. Reise Mossambique 6: 549. 1863. MOZAMBIQUE: Tete, s.d., Peters s.n. (holotype: B).
Echinochloa equitans Echinochloa equitans (A. Richard) C. E. Hubbard, in Troupin, Expl. Parc Nat. Garamba 4: 33. 1956.Panicum equitans A. Richard 1850. 
Echinochloa verticillata Echinochloa verticillata Berhaut, Mém. Soc. Bot. France 1953-1954: 9. 1954 SENEGAL
Echinochloa zonalis Echinochloa zonalis (Gussone) Parlatore, Fl. Panorm. 119. 1839.Panicum zonale Gussone 1827. 
Milium colonum Milium colonum (Linnaeus) Moench, Methodus 202. 1794.Panicum colonum Linnaeus 1759. 
Oplismenus colonus Oplismenus colonus (Linnaeus) Kunth, in Humboldt et al., Nov. Gen. Sp. 1: 108. 1816.Panicum colonum Linnaeus 1759. 
Oplismenus colonus var. zonalis Oplismenus colonus (Linnaeus) Kunth, var. zonalis (Gussone) Schrader, Linnaea 12: 429. 1838.Panicum zonale Gussone 1827. 
Oplismenus crus-galli var. colonus Oplismenus crus-galli (Linnaeus) Dumortier, var. colonus (Linnaeus) Cosson, in Durieu de Maisonnueve, Expl. Sci. Algérie 2: 28. 1855 ("1849").Panicum colonum Linnaeus 1759. 
Oplismenus daltonii Oplismenus daltonii (Parlatore ex Hooker) J. A. Schmidt, Beitr. Fl. Cap. Verd. Ins. 136. 1852.Panicum daltonii Parlatore ex Hooker 1849. 
Oplismenus muticus Oplismenus muticus Philippi, Anales Univ. Chile 93: 714. 1896. CHILE
Oplismenus repens Oplismenus repens J. Presl, in C. Presl, Reliq. Haenk. 1: 321. 1830. MEXICO: Without data, Haenke s.n. (holotype: PR).
Oplismenus pseudocolonus Oplismenus pseudocolonus (Roth ex Roemer & Schultes) Kunth, Revis. Gramin. 44. 1829.Panicum pseudocolonum Roth ex Roemer & Schultes 1817. 
Panicum colonum var. angustatum Panicum colonum Linnaeus, var. angustatum Peter, Repert. Spec. Nov. Regni Veg. Beih. 40(1, Anh.): 33. 1930. TANZANIA: Kigoma Distr.: Uvinza, s.d., Peter 35953 (holotype: B(destroyed)).
Panicum colonum var. atroviolaceum Panicum colonum Linnaeus, var. atroviolaceum Hackel, Vierteljahrsschr. Naturf. Ges. Zürich 56: 71. 1911. SOUTH-WEST AFRICA: Namibia: Ondonga to Uakuyanyama, s.d., Rautanen 741 (holotype: Z).
Panicum colonum var. equitans Panicum colonum Linnaeus, var. equitans (A. Richard) T. Durand & Schinz, Consp. Fl. Afric. 5: 743. 1894.Panicum equitans A. Richard 1850. 
Panicum colonum var. humile Panicum colonum Linnaeus, var. humile Nees von Esenbeck, Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(Suppl. 1): 139. 1843. PERU
Panicum colonum var. zonale Panicum colonum Linnaeus, var. zonale (Gussone) L. H. Dewey, in J. M. Coulter, Contr. U.S. Natl. Herb. 2: 502. 1894.Panicum zonale Gussone 1827. 
Panicum crus-galli subsp. colonum Panicum crus-galli Linnaeus, subsp. colonum (Linnaeus) Makino & Nemoto, Fl. Japan 1470. 1925.Panicum colonum Linnaeus 1759. 
Panicum crusgalli var. colonum Panicum crusgalli Linnaeus, var. colonum (Linnaeus) Cosson ex Richter, Pl. Eur. 1: 26. 1890.Panicum colonum Linnaeus 1759. 
Panicum crusgalli var. longisetum Panicum crusgalli Linnaeus, var. longisetum Trinius, Sp. Gram. 2: t. 162. 1829.  
Panicum daltonii Panicum daltonii Parlatore ex Hooker, Niger Fl. 185. 1849. CAPE VERDE
Panicum echinochloa Panicum echinochloa T. Durand & Schinz, Consp. Fl. Afric. 5: 748. 1894.Echinochloa divaricata Andersson 1863. 
Panicum equitans Panicum equitans A. Richard, Tent. Fl. Abyss. 2: 365. 1850. ETHIOPIA: Sara, s.d., Schimper 1068 (holotype: ?; isotype: K).
Panicum hookeri Panicum hookeri Parlatore, Atti Congr. Sci. Ital. 8: 586. 1847. CAPE VERDE
Panicum prorepens Panicum prorepens Steudel, Syn. Pl. Glumac. 1: 46. 1853.Oplismenus repens J. Presl 1830. 
Panicum tetrastichum Panicum tetrastichum Forsskål, Fl. Aegypt.-Arab. 19. 1775. EGYPT
Panicum zonale Panicum zonale Gussone, Fl. Sicul. Prodr. 1: 62. 1827. ITALY: Sicily
Panicum brachiariiforme Panicum brachiariiforme Steudel, Syn. Pl. Glumac. 1: 54. 1853. West Africa, without data, Le Jolis s.n. (holotype: P).
Panicum pseudocolonum Panicum pseudocolonum Roth ex Roemer & Schultes, Syst. Veg. 2: 424. 1817.  
Setaria brachiariiformis Setaria brachiariiformis (Steudel) T. Durand & Schinz, Consp. Fl. Afric. 5: 772. 1894.Panicum brachiariiforme Steudel 1854 
Panicum colonum Panicum colonum Linnaeus, Syst. Nat., ed. 10. 870. 1759. JAMAICA: Without data, Browne s.n. (lectotype: LINN 80.23). Lectotypified by Clayton & Renvoize, in Polhill, Fl. Trop. E. Afr., Gramin. 557. 1982.

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Blount, Bullock, Butler, Calhoun, Chilton, Conecuh, Coosa, Crenshaw, Dallas, Elmore, Escambia, Etowah, Franklin, Greene, Hale, Houston, Lamar, Lawrence, Lee, Limestone, Lowndes, Madison, Marengo, Marshall, Mobile, Morgan, Pike, Tuscaloosa, Wilcox

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: AUA, JSU, TROY, UNA, UWAL, UWFP

Range of years during which specimens were collected: 1957 - 1992

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
UWAL0043040 Baldwin 10 Aug 2002 Larsen, H.S.
1070
UWAL
Echinochloa colona (L.) Link
UNA00050662 Marshall 02 Aug 1992 Spaulding, Daniel D.
2547
UNA
Echinochloa colona (L.) Link
UWAL0019197 Chilton 07 Jul 1990 Kral, R.
77870
UWAL
Echinochloa colonum (L.) Link
AUA000024460 Baldwin 19 JUL 2007 SPAULDING, DANIEL D.
12723
AUA
ECHINOCHLOA COLONA (L.) LINK
UNA00003713 Mobile s.d. Mohr, Charles
s.n.
UNA
Echinochloa colona (L.) Link
UWAL0002341 Greene 07 Aug 2008 Spaulding, Daniel D.
13047
UWAL
Echinochloa colona (L.) Link
UWAL0032924 Morgan 15 Aug 2015 England, J. Kevin
5605
UWAL
Echinochloa colonum (Linnaeus) Link
TROY000044561 Greene 07 Aug 2008 Spaulding, Daniel D.
13047
TROY
Echinochloa colonum (Linnaeus) Link
UWAL0046562 Autauga 21 Sep 2017 Parker, Matthew R.
1302
UWAL
Echinochla colonum (Linnaeus) Link
JSU101115 Franklin 15 Oct 1975 Whetstone, R. David
7064
JSU
Echinochloa colona (Linnaeus) Link
TROY000049241 Autauga 14 Sep 2018 Parker, Matthew R.
1780
TROY
Echinochloa colona (Linnaeus) Link
UWAL0038832 Bullock 30 Sep 2017 Keener, Brian R.
4293
UWAL
Echinochloa colona (Linnaeus) Link
UNA00058654 Lamar 28 Jul 2002 Keener, Brian R.
2317
UNA
Echinochloa colona (L.) Link
UNA00062298 Dallas 28 Jun 2003 Spaulding, Daniel D.
11927
UNA
Echinochloa colona (L.) Link
TROY000045240 Autauga 21 Sep 2017 Parker, Matthew R.
1302
TROY
Echinochla colonum (Linnaeus) Link
UWAL0035759 Mobile 01 Aug 2015 Horne, Howard E.
3004
UWAL
Echinochloa colonum (Linnaeus) Link
UWFP000020805acc Pike 06 Oct 2006 James R. Burkhalter
21173
UWFP
Echinochloa colona (Linnaeus) Link
AUA000016935 Baldwin JONES, JR., SAMUEL B.
308
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000047092 Baldwin 13 Jul 1974 Lelong, Michel G.
8003
TROY
Echinochloa colona (Linnaeus) Link
AUA000016947 Lowndes 8 NOV 1992 DIAMOND, JR., ALVIN R.
8502
AUA
ECHINOCHLOA COLONA (L.) LINK
JSU101113 Franklin 15 Oct 1975 Whetstone, R. David
7064
JSU
Echinochloa colona (Linnaeus) Link
TROY000028297 Marengo 20 Aug 2008 England, J. Kevin
913
TROY
Echinochloa colona (Linnaeus) Link
UWAL0038407 Mobile 17 Aug 1974 LeLong, Michel G.
8154
UWAL
Echinochloa colona (Linnaeus) Link
UNA00003716 Tuscaloosa 01 Aug 1957 Deason, Temd R.
s.n.
UNA
Echinochloa colona (L.) Link
TROY000041178 Elmore 20 Apr 2012 Pullen, Robert
418
TROY
Echinochloa colona (Linnaeus) Link
AUA000016937 Conecuh 19 AUG 1995 DIAMOND, JR., ALVIN R.
9866
AUA
ECHINOCHLOA COLONA (L.) LINK
AUA000016936 Chilton 27 JUL 2005 DIAMOND, JR., ALVIN R.
15551
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000012131 Houston 08 Sep 2002 Spaulding, Daniel D.
11582
TROY
Echinochloa colona (Linnaeus) Link
AUA000016943 Lee 29 SEP 1971 BOONE, ALLISON
056
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000048022 Lawrence 02 Oct 1993 Blythe, Fennel
25
TROY
Echinochloa colona (Linnaeus) Link
AUA000016938 Crenshaw 31 JUL 1994 DIAMOND, JR., ALVIN R.
9275
AUA
ECHINOCHLOA COLONA (L.) LINK
JSU101112 Etowah 27 Aug 1994 Hodge, Lesley M.
3242
JSU
Echinochloa colona (Linnaeus) Link
TROY000026783 Barbour 17 Jul 2008 Diamond, Alvin R.
19559
TROY
Echinochloa colona (Linnaeus) Link
AUA000016945 Lee 17 AUG 1966 HARRIS, JOHN
AUA
ECHINOCHLOA COLONA (L.) LINK
UNA00059489 Houston 8 Sep 2002 Spaulding, Daniel D.
11582
UNA
Echinochloa colona (L.) Link
AUA000016948 Mobile 14 AUG 1996 ANDERSON, LORAN C.
16,862
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000012132 Wilcox 09 Oct 2006 Diamond, Alvin R.
17075
TROY
Echinochloa colona (Linnaeus) Link
UWAL0039743 Houston 18 Aug 2016 Diamond, Alvin R.
27615
UWAL
Echinochloa colona (Linnaeus) Link
TROY000012129 Butler 06 Jul 2007 Diamond, Alvin R.
18037
TROY
Echinochloa colona (Linnaeus) Link
UWAL0036578 Mobile 23 Sep 2017 Horne, Howard E.
4235
UWAL
Echinochloa colona (Linnaeus) Link
AUA000038120 Morgan 15 Aug 2015 ENGLAND, J. KEVIN
5605
AUA
ECHINOCHLOA COLONUM (L.) LINK
AUA000016949 Mobile 31 AUG 1960 JONES, JR., SAMUEL B.
307
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000047258 Baldwin 20 Oct 1975 Lelong, Michel G.
8860
TROY
Echinochloa colona (Linnaeus) Link
UWAL0032023 Houston 13 Aug 2015 Diamond, Alvin R.
26738
UWAL
Echinochloa colona (Linnaeus) Link
JSU101114 Franklin 15 Oct 1975 Whetstone, R. David
7064
JSU
Echinochloa colona (Linnaeus) Link
TROY000047148 Mobile 17 Aug 1974 Lelong, Michel G.
8154
TROY
Echinochloa colona (Linnaeus) Link
UNA00069245 Marengo 20 Aug 2008 England, J. Kevin
913
UNA
Echinochloa colona (L.) Link
UNA00003733 Tuscaloosa 01 Aug 1957 Deason, Temd R.
s.n.
UNA
Echinochloa colona (L.) Link
TROY000027128 Chilton 27 Jul 2005 Diamond, Alvin R.
15551
TROY
Echinochloa colona (Linnaeus) Link
UNA00003787 Hale 30 Jun 1966 Maginness, Sherrie E.
341
UNA
Echinochloa colona (L.) Link
UWAL0036031 Blount 28 Jul 2013 Anderson, Kelly
0412
UWAL
Echinochloa colonum (L.) Link
TROY000012130 Dallas 28 Sep 2003 Spaulding, Daniel D.
11927
TROY
Echinochloa colona (Linnaeus) Link
UWFP000023351acc Escambia 04 Sep 2010 James R. Burkhalter
s.n.
UWFP
Echinochloa colona (Linnaeus) Link
AUA000016946 Lee 24 JUL 1969 JOHNSON, W. C.
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000012126 Barbour 26 Jul 2000 Diamond, Alvin R.
11903
TROY
Echinochloa colona (Linnaeus) Link
AUA000016939 Crenshaw 7 AUG 1995 DIAMOND, JR., ALVIN R.
9812
AUA
ECHINOCHLOA COLONA (L.) LINK
JSU101117 Franklin 15 Sep 1996 Spaulding, Daniel D.
9224
JSU
Echinochloa colona (Linnaeus) Link
TROY000012128 Butler 16 Aug 2007 Diamond, Alvin R.
17921
TROY
Echinochloa colona (Linnaeus) Link
UWAL0052039 Madison 04 Oct 2020 England, J. Kevin
11351
UWAL
Echinochloa colonum (L.) Link
AUA000016950 Pike 16 AUG 1995 DIAMOND, JR., ALVIN R.
9850
AUA
ECHINOCHLOA COLONA (L.) LINK
UWAL0008961 Butler 06 Jul 2007 Diamond, A.R.
18037
UWAL
Echinochloa colona (L.) Link
TROY000044148 Houston 18 Aug 2016 Diamond, Alvin R.
27615
TROY
Echinochloa colona (Linnaeus) Link
AUA000016941 Elmore 18 JUL 1967 TEEM, DAVID H.
17
AUA
ECHINOCHLOA COLONA (L.) LINK
JSU101111 Calhoun 24 Aug 1982 Whetstone, R. David
11979
JSU
Echinochloa colona (Linnaeus) Link
UWAL0008960 Baldwin 19 Jul 2007 Spaulding, Daniel D.
12723
UWAL
Echinochloa colona (L.) Link
UNA00003711 Tuscaloosa 30 Aug 1961 Harper, Roland M.
4483
UNA
Echinochloa colona (L.) Link
UNA00003701 Mobile 15 Aug 1966 Deramus, Rebecca
D998
UNA
Echinochloa colona (L.) Link
JSU101116 Franklin 15 Oct 1975 Whetstone, R. David
7064
JSU
Echinochloa colona (Linnaeus) Link
UWAL0043039 Baldwin 25 Sep 2001 Larsen, H.S.
916
UWAL
Echinochloa colona (L.) Link
TROY000026871 Baldwin 19 Jul 2007 Spaulding, Daniel D.
12723
TROY
Echinochloa colona (Linnaeus) Link
AUA000016940 Elmore 5 AUG 1969 REBOIS, R. V.
155
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000027129 Lamar 28 Jul 2002 Keener, Brian R.
2317
TROY
Echinochloa colona (Linnaeus) Link
UWAL0038406 Baldwin 13 Jul 1974 LeLong, Michel G.
8003
UWAL
Echinochloa colona (Linnaeus) Link
AUA000016952 Pike 17 OCT 1994 DIAMOND, JR., ALVIN R.
9417
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000027127 Blount 24 May 1998 Keener, Brian R.
1291
TROY
Echinochloa colona (Linnaeus) Link
TROY000042777 Houston 13 Aug 2015 Diamond, Alvin R.
26738
TROY
Echinochloa colona (Linnaeus) Link
UWAL0014860 Coosa 12 Jul 2013 Diamond, Alvin R.
24341
UWAL
Echinochloa colona (Linnaeus) Link
UNA00053535 Greene 5 Aug 1969 Thomas, Joab L.
2383
UNA
Echinochloa colona (L.) Link
AUA000016951 Pike 3 AUG 1994 DIAMOND, JR., ALVIN R.
9278
AUA
ECHINOCHLOA COLONA (L.) LINK
UNA00003706 Hale 25 Sep 1965 Williams, Norris H.
133
UNA
Echinochloa colona (L.) Link
TROY000012127 Butler 02 Sep 2007 Diamond, Alvin R.
18019
TROY
Echinochloa colona (Linnaeus) Link
AUA000024838 Escambia 8 JUL 2009 HANSEN, CURTIS J.
4091
AUA
ECHINOCHLOA COLONUM (L.) LINK
AUA000016942 Lee 27 JUN 1972 NELSON, JOHN
047
AUA
ECHINOCHLOA COLONA (L.) LINK
TROY000036692 Bullock 26 Jul 2012 Dykes, Bailee
1080
TROY
Echinochloa colona (Linnaeus) Link
AUA000016934 Bullock 17 AUG 1993 DIAMOND, JR., ALVIN R.
8739
AUA
ECHINOCHLOA COLONA (L.) LINK
AUA000016944 Lee 16 JUL 1969 WEAVER, JIMMY
077
AUA
ECHINOCHLOA COLONA (L.) LINK
JSU101110 Calhoun 24 Aug 1982 Whetstone, R. David
11979
JSU
Echinochloa colona (Linnaeus) Link
TROY000037563 Coosa 12 Jul 2013 Diamond, Alvin R.
24341
TROY
Echinochloa colona (Linnaeus) Link
UNA00058553 Blount 24 May 1998 Keener, Brian R.
1291
UNA
Echinochloa colona (L.) Link
UWAL0014679 Bullock 26 Jul 2012 Dykes, Bailee
1080
UWAL
Echinochloa colona (Linnaeus) Link
UNA00003792 Limestone 01 Aug 1978 Haynes, Robert R.
6804
UNA
Echinochloa colona (L.) Link
TROY000026784 Lowndes 02 Nov 2008 Diamond, Alvin R.
20337
TROY
Echinochloa colona (Linnaeus) Link
UWAL0008962 Marengo 20 Aug 2008 England, J. Kevin
0913
UWAL
Echinochloa colona (L.) Link
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