Ilex opaca

Family:AQUIFOLIACEAE
Species:Ilex opaca Aiton
Common Name:American Holly
Habitat:Mesic to dry forests, alluvial forests, sandhills, and thickets.
Associated Ecological Communities:**
Growth Habit:Tree
Duration:Perennial
Category:Vascular
USDA Symbol:ILOP
Plant Notes:American Holly is a native evergreen tree in the Holly family (Aquifoliaceae). It occurs statewide in Alabama. American Holly grows in a variety of forest types from disturbed woodlands to mature hardwood forest, and on floodplains. This small to medium sized tree grows between 30 and 50 feet in height. The bark is smooth and light gray to tan in color. The leaves are alternate, petiolate, evergreen, elliptic in outline with sharp spine tipped teeth along their margins. The leaves are thick and leathery in texture. American Holly is dioecious, with separate male and female plants. The flowers are small and greenish white. Male flowers occur in axillary cymes of 3-7 flowers. The flowers have 4 green sepals, 4 greenish-white petals, and 4 stamens, and no pistil. The female flowers are solitary or in axillary cymes of 1-3. Female flowers have 4 green sepals, 4 greenish-white petals, 4 sterile stamens, and a single pistil. The fruit (only produced by the female plant) is a bright red or occasionally orange or yellow drupe. Each fruit contains 4 hard seed (pyrenes). The seed are tan in color and irregularly angled on the back. Many species of birds and small mammals feed on the fruit. American Holly is a popular small tree for the home landscape. It tolerates a wide range of soil and moisture types. It is also shade tolerant and not browsed by deer. Sprigs of American Holly are often used as Christmas decorations. The fruit are slightly toxic to humans, and should not be eaten. Many cultivars are available including ones with yellow fruit, dwarf forms, and ones with spineless leaves. There are also hybrids between the American Holly and other species. Ilex ×attenuata Ashe (often called the “Savannah Holly”) is a hybrid between American Holly and Dahoon Holly (Ilex cassine Linnaeus).--A. Diamond
Taxonomic Notes:**
Status:Native, FAC- (NWI)
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyAQUIFOLIACEAE - Holly family
Genus Ilex
Species Ilex opaca Aiton - American Holly

Citation

Citation Ilex opaca Aiton, Hort. Kew. 1: 169. 1789.
Basionym: **
Type: **

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Ageria opaca Ageria opaca (Aiton) Rafinesque, Sylva Tellur. 47. 1838.Ilex opaca Aiton 1789. 
Ilex laxiflora Ilex laxiflora Lamarck, Encycl. 3: 147. 1789.  
Ilex opaca var. latifolia Ilex opaca Aiton, var. latifolia Beissner, in Beissner et al., Handb. Laubholzben. 290. 1903.  
Ilex opaca var. laxiflora Ilex opaca Aiton, var. laxiflora (Lamarck) Nuttall, Gen. N. Amer. Pl. 1: 109. 1818.Ilex laxiflora Lamarck 1789. 
Ilex opaca var. macrodon Ilex opaca Aiton, var. macrodon Beissner, in Beissner et al., Handb. Laubholzben. 290. 1903.  
Ilex opaca var. subintegra Ilex opaca Aiton, var. subintegra (Weatherby) Rehder, Man. Cult. Trees 543. 1927.Ilex opaca Aiton, forma subintegra Weatherby 1921. 
Ilex opaca var. globosa Ilex opaca Aiton, var. globosa Beissner, in Beissner et al., Handb. Laubholzben. 290. 1903.  
Ilex opaca var. opaca Ilex opaca Aiton, var. integra A. W. Wood, Amer. Bot. Fl. 207. 1870.  
Ilex opaca var. acuminata Ilex opaca Aiton, var. acuminata Beissner, in Beissner et al., Handb. Laubholzben. 290. 1903.  
Ilex opaca var. floribunda Ilex opaca Aiton, var. floribunda Demcker, Mitt. Deutsch. Dendrol. Ges. 1904: 155. 1904. NEW YORK
Ilex opaca var. xanthocarpa Ilex opaca Aiton, var. xanthocarpa Rehder, Mitt. Deutsch. Dendrol. Ges. 1907: 73. 1907.  
Ilex quercifolia Ilex quercifolia Meerburgh, Pl. Select. Icon. Pict. t. 5. 1798.  

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Bibb, Blount, Bullock, Butler, Calhoun, Chambers, Cherokee, Chilton, Choctaw, Clarke, Clay, Cleburne, Coffee, Colbert, Conecuh, Covington, Crenshaw, Cullman, Dale, Dallas, DeKalb, Elmore, Escambia, Etowah, Franklin, Geneva, Hale, Henry, Houston, Jackson, Lamar, Lauderdale, Lawrence, Lee, Limestone, Lowndes, Macon, Madison, Marengo, Marion, Marshall, Mobile, Monroe, Montgomery, Morgan, Perry, Pike, Randolph, St. Clair, Sumter, Talladega, Tallapoosa, Tuscaloosa, Walker, Washington, Wilcox, Winston

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AMAL, AUA, BRIT, JSU, NY, SAMF, TROY, UNA, UNAF, USAM, UWAL

Range of years during which specimens were collected: 1895 - 2019

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
UNAF0000516 Lauderdale 02 Sep 1989 den Boer, John H.
s.n.
UNAF
Ilex opaca Aiton
UNAF0000523 Marion 03 Sep 1989 Markham, Todd
s.n.
UNAF
Ilex opaca Aiton
JSU119418 Cleburne 28 Sep 1982 Bussey, Melanie G.
469
JSU
Ilex opaca Aiton
AUA000032190 Tallapoosa 17 Apr 1973 Cannon, Ed
108
AUA
Ilex Opaca Ait.
AUA000031869 Barbour 11 Oct 1975 Wallace, Chip
056
AUA
Ilex Opaca Ait.
TROY000033679 Washington 16 Apr 2011 Keener, Brian R.
6364
TROY
Ilex opaca Aiton
USAM000001426 Mobile 28 Nov 1970 Rush, M.C.
s.n.
USAM
Ilex opaca Aiton
JSU119621 Randolph 13 Jul 1969 Herren, Tommy
s.n.
JSU
Ilex opaca Aiton
AUA000031873 Butler 25 Apr 1986 Diamond, Alvin R.
2345
AUA
Ilex Opaca Ait.
AUA000031884 Covington 19 Jun 2008 Hansen, Curtis J.
3265
AUA
Ilex Opaca Ait. Var. Opaca
UNA00073940 Mobile 19 Mar 2017 Horne, Howard
3742
UNA
Ilex opaca Ait.
UWAL0056481 Bibb 11 Jun 2019 Keener, Brian R.
11359
UWAL
Ilex opaca Aiton
TROY000045715 Autauga 11 Nov 2016 Parker, Matthew R.
302
TROY
Ilex opaca Aiton
SAMF0003209 Bibb 10 Jun 2015 Davenport, Lawrence J.
5879
SAMF
Ilex opaca Aiton
AMAL00016070 Barbour 05 Jan 2006 Diamond, Alvin R.
16112
AMAL
Ilex opaca Ait.
AUA000031910 Macon 21 Jul 1972 Hallmark, Wendy
087
AUA
Ilex Opaca Ait.
UWAL0012068 Conecuh 28 Jun 2011 Keener, Brian R.
6573
UWAL
Ilex opaca Aiton
TROY000032565 Houston 01 May 2011 Diamond, Alvin R.
22022
TROY
Ilex opaca Aiton
BRIT73411 DeKalb 25 Jul 1970 Holcombe, W.H.
86
BRIT
Ilex opaca Aiton
JSU119398 Calhoun 29 Jun 1988 Landers, Kenneth E.
s.n.
JSU
Ilex opaca Aiton
UNAF0000520 Lawrence 15 Sep 1989 Crews, Sherry
s.n.
UNAF
Ilex opaca Aiton
UNAF0000512 Franklin 06 Oct 1985 Jackson, Carle
s.n.
UNAF
Ilex opaca Aiton
JSU119419 Cleburne 20 Oct 1982 Bussey, Melanie G.
559
JSU
Ilex opaca Aiton
AUA000031906 Lee 6 May 1959 Brady, Jr., V.E.
AUA
Ilex Opaca Ait.
AUA000032189 Tallapoosa 11 Apr 1997 Tyler, Patricia J.
TYLER14
AUA
Ilex Opaca Ait.
TROY000044332 Monroe 16 Jan 2017 Diamond, Alvin R.
27759
TROY
Ilex opaca Aiton
JSU119415 Cleburne 3 Oct 1972 Guinn, Sara
s.n.
JSU
Ilex opaca Aiton
AMAL00017726 Lauderdale 25 Jun 2009 Holt, Brian D.
SC-85
AMAL
Ilex opaca Ait.
AUA000031878 Coffee 27 Jul 1989 Diamond, Alvin R.
6170
AUA
Ilex Opaca Ait.
UWAL0000306 Lamar 13 Aug 2002 Keener, Brian R.
2401
UWAL
Ilex opaca Aiton
UNA00068708 Lamar 13 Aug 2002 Keener, Brian R.
2401
UNA
Ilex opaca Ait.
TROY000001181 Monroe 29 Dec 2005 Diamond, Alvin R.
16110
TROY
Ilex opaca Aiton
JSU119427 Franklin 17 Mar 1968 South, Jane
s.n.
JSU
Ilex opaca Aiton
AUA000031896 Hale 12 Nov 1993 Diamond, Alvin R.
8978
AUA
Ilex Opaca Ait.
UWAL0002631 Walker 21 Dec 2010 England, J. Kevin
2629
UWAL
Ilex opaca Aiton
SAMF0008286 Covington 12 Apr 2009 Dansby, Elliott
25
SAMF
Ilex opaca Aiton
BRIT73406 Covington 20 Mar 1973 Gonsoulin, G.
1937
BRIT
Ilex opaca Aiton
JSU119420 Colbert 16 Mar 1967 Landers, Kenneth E.
s.n.
JSU
Ilex opaca Aiton
AUA000032185 Montgomery 15 Jan 1989 Diamond, Alvin R.
5647
AUA
Ilex Opaca Ait.
UWAL0035586 Mobile 04 Jul 2014 Horne, Howard E.
2621
UWAL
Ilex opaca Aiton
UNAF0000511 Franklin 07 Sep 1990 Davidson, Nathan
s.n.
UNAF
Ilex opaca Aiton
UNAF0000510 Crenshaw 17 Jun 1991 Lovelace, Connie
s.n.
UNAF
Ilex opaca Aiton
JSU119417 Cleburne 28 Sep 1982 Bussey, Melanie G.
469
JSU
Ilex opaca Aiton
AUA000032188 Tallapoosa 15 May 1965 Brown, Jimmy E.
84
AUA
Ilex Opaca Ait.
AUA000032184 Monroe 18 Apr 2003 Valente, M.J.
44
AUA
Ilex Opaca Ait.
TROY000014264 Cherokee 26 Sep 1987 Dickson, Catherine C.
694
TROY
Ilex opaca Aiton
SAMF0005110 DeKalb 14 May 2006 Davenport, Lawrence J.
3428
SAMF
Ilex opaca Aiton
JSU119423 DeKalb 15 Mar 1955 Hanson, Gene
s.n.
JSU
Ilex opaca Aiton
AMAL00005825 Marion 03 Jun 2002 Keener, Brian R.
2163
AMAL
Ilex opaca Ait.
AUA000031871 Bibb 27 Jul 2003 Diamond, Alvin R.
14042
AUA
Ilex Opaca Ait.
UNA00073799 Washington 27 Jul 2012 England, J. Kevin
3810
UNA
Ilex opaca Ait.
UNA00020687 Tuscaloosa 10 May 1968 Thomas, Joab L.
1682
UNA
Ilex opaca Ait.
TROY000040741 Conecuh 28 Jun 2011 Keener, Brian R.
6573
TROY
Ilex opaca Aiton
AMAL00005818 Conecuh 05 Jan 2002 Diamond, Alvin R.
12849
AMAL
Ilex opaca Ait.
AUA000031875 Cherokee 30 Apr 1972 Hale, A.R.
109
AUA
Ilex Opaca Ait.
UWAL0000307 Marengo 24 Sep 2008 England, J. Kevin
1180
UWAL
Ilex opaca Aiton
TROY000029849 Barbour 01 Oct 2004 Dasinger, Keith
4
TROY
Ilex opaca Aiton
BRIT73424 Madison 09 May 1964 Heller, H.
92
BRIT
Ilex opaca Aiton
JSU143078 Colbert 16 Mar 1967 Landers, Kenneth E.
s.n.
JSU
Ilex opaca Aiton
AUA000031883 Conecuh 16 Dec 1985 Diamond, Alvin R.
1960
AUA
Ilex Opaca Ait.
UNAF0000514 Jackson 01 Sep 1989 Gamble, Michael
s.n.
UNAF
Ilex opaca Aiton
USAM000001434 Mobile 24 Jul 1969 LeLong, Michel G.
5186
USAM
Ilex opaca Aiton
JSU119433 Monroe 1 Nov 1997 Whetstone, R. David
19093
JSU
Ilex opaca Aiton
AUA000031899 Lee 25 May 1971 Fuller, C.W.
106
AUA
Ilex Opaca Ait.
AUA000032178 Monroe 18 Oct 1980 Webb, A.L.
131
AUA
Ilex Opaca Ait.
USAM000001435 Mobile 18 Mar 1969 Wofford, B. Eugene
262
USAM
Ilex opaca Aiton
JSU119400 Calhoun 1 Nov 1992 Spaulding, Daniel D.
3066
JSU
Ilex opaca Aiton
AUA000031880 Conecuh 26 Apr 1986 Diamond, Alvin R.
2382
AUA
Ilex Opaca Ait.
UWAL0017634 Houston 02 Apr 2015 Diamond, Alvin R.
26115
UWAL
Ilex opaca Aiton
UNA00050253 Cherokee 30 Oct 1986 Moore, C.M.
100
UNA
Ilex opaca Ait.
ALNHS00001555 Clarke 17 Feb 2012 Horne, Howard E.
SWS-12
ALNHS
Ilex opaca Ait.
BRIT73398 Clarke 24 Jun 1984 Whetstone, R. David
14310
BRIT
Ilex opaca Aiton
JSU119628 Winston 13 Aug 1974 Atkinson, Tim A.
710
JSU
Ilex opaca Aiton
AMAL00005822 Escambia 10 Feb 2002 Diamond, Alvin R.
12897
AMAL
Ilex opaca Ait.
AUA000031889 DeKalb 25 Jun 1977 Quenelle, M.A.
25
AUA
Ilex Opaca Ait.
UNA00058063 Marion 03 Jun 2002 Keener, Brian R.
2163
UNA
Ilex opaca Ait.
TROY000001186 Pike 27 Oct 1995 Holmes, Jeanese
106
TROY
Ilex opaca Aiton
SAMF0005108 Washington 16 Apr 2011 Keener, Brian R.
6364
SAMF
Ilex opaca Aiton
JSU144265 Cherokee 31 May 1968 Landers, Kenneth E.
s.n.
JSU
Ilex opaca Aiton
AUA000031916 Macon 21 Jul 1972 Nelson, John
083
AUA
Ilex Opaca Ait.
UNAF0000526 Winston 07 Jul 1990 Beasley, James
s.n.
UNAF
Ilex opaca Aiton
UNAF0000517 Lauderdale 10 Oct 1989 Grosso, Vincent
s.n.
UNAF
Ilex opaca Aiton
USAM000001436 27 Apr 1975 LeLong, Michel G.
8424
USAM
Ilex opaca Aiton
AMAL00005824 Madison 13 Aug 1996 Threlkeld, Steven J.
823
AMAL
Ilex opaca Ait.
AUA000031898 Lee 14 Apr 1936 Thompson, C.J.
AUA
Ilex Opaca Ait.
AUA000031918 Marengo 7 Nov 1982 Wilburn, John C.
153
AUA
Ilex Opaca Ait.
JSU143076 Calhoun 18 May 1971 Edmondson, Fay
s.n.
JSU
Ilex opaca Aiton
JSU119421 Cullman 18 Mar 1994 Whetstone, R. David
s.n.
JSU
Ilex opaca Aiton
AUA000032193 Wilcox 26 Apr 1986 Diamond, Alvin R.
2412
AUA
Ilex Opaca Ait.
UWAL0011520 Crenshaw 02 Sep 2012 Diamond, Alvin R.
23292
UWAL
Ilex opaca Aiton
UNA00050252 Randolph 11 Oct 1985 Nixon, C.F.
2017
UNA
Ilex opaca Ait.
TROY000001187 Wilcox 07 Dec 2005 Diamond, Alvin R.
16083
TROY
Ilex opaca Aiton
JSU119426 Etowah 23 Apr 1994 Hodge, Lesley M.
1190
JSU
Ilex opaca Aiton
AMAL00005821 Elmore 13 Jan 2002 Diamond, Alvin R.
12856
AMAL
Ilex opaca Ait.
AUA000031894 Escambia 3 Jul 1970 Rogers, Norman
069
AUA
Ilex Opaca Ait.
UNA00050895 Marion 01 Jul 1985 McDaniel, Sidney
28221
UNA
Ilex opaca Ait.
TROY000001183 Pike 13 Apr 1991 Huggins, Gwyn
19
TROY
Ilex opaca Aiton
USAM000001429 Baldwin 20 Apr 1974 LeLong, Michel G.
7616
USAM
Ilex opaca Aiton
JSU119402 Cherokee 8 Jul 1969 Tudor, Jessica
s.n.
JSU
Ilex opaca Aiton
UWAL0002822 Covington 20 Dec 2010 Diamond, Alvin R.
21722
UWAL
Ilex opaca Aiton
UNAF0000525 Morgan 04 Jul 1991 Engle, Dennis
s.n.
UNAF
Ilex opaca Aiton
UNAF0000519 Lauderdale 17 Oct 1983 McGregor, Stuart
s.n.
UNAF
Ilex opaca Aiton
JSU119397 Calhoun 7 Mar 1987 Lawler, Carol L.
1500
JSU
Ilex opaca Aiton
AUA000031917 Macon 21 Jul 1972 Eudaly, Edwin M.
114
AUA
Ilex Opaca Ait.
AUA000031912 Macon 9 May 1953 Walsh, C.S.
AUA
Ilex Opaca Ait.
USAM000001430 Mobile 18 Mar 1969 Wofford, B. Eugene
262
USAM
Ilex opaca Aiton
JSU119407 Choctaw 13 Sep 1997 Higginbotham, Jerri W.
247
JSU
Ilex opaca Aiton
AUA000031882 Conecuh 25 Apr 1986 Diamond, Alvin R.
2365
AUA
Ilex Opaca Ait.
UWAL0005369 Houston 01 May 2011 Diamond, Alvin R.
22022
UWAL
Ilex opaca Aiton
UNA00073382 Lee 22 Oct 1996 Miller, James H.
s.n.
UNA
Ilex opaca Ait.
ALNHS00004369 Houston 01 Nov 2016 Barger, T. Wayne
SP#2731
ALNHS
Ilex opaca Ait.
TROY000047790 Autauga 14 Dec 2017 Parker, Matthew R.
1486
TROY
Ilex opaca Aiton
USAM000001431 Clarke 13 Jul 1969 Kral, Robert
35485
USAM
Ilex opaca Aiton
AMAL00016074 Wilcox 07 Dec 2005 Diamond, Alvin R.
16083
AMAL
Ilex opaca Ait.
AUA000031909 Macon 22 Jul 1972 Pounders, C.T.
119
AUA
Ilex Opaca Ait.
UWAL0036707 Lamar 13 Jul 2017 England, J. Kevin
8127
UWAL
Ilex opaca Aiton
TROY000014267 Elmore 13 Jan 2002 Diamond, Alvin R.
12856
TROY
Ilex opaca Aiton
BRIT73415 Escambia 21 Apr 1962 Ingram, D.
97
BRIT
Ilex opaca Aiton
JSU119404 Cherokee 31 May 1968 Landers, Kenneth E.
s.n.
JSU
Ilex opaca Aiton
AUA000031891 Escambia 23 Oct 1982 Graves, Tom
85
AUA
Ilex Opaca Ait.
UNA00056414 Dallas 21 Sep 1995 Birk, Michelle
419
UNA
Ilex opaca Ait.
UNAF0000521 Limestone 11 Aug 1989 Butler, Yvette
s.n.
UNAF
Ilex opaca Aiton
UNAF0000518 Lauderdale 02 Sep 1989 Hardy, Dawn
s.n.
UNAF
Ilex opaca Aiton
USAM000001428 Monroe 27 Apr 1975 LeLong, Michel G.
8442
USAM
Ilex opaca Aiton
AUA000031904 Lee 11 Apr 1969 Barnes, W.K.
043
AUA
Ilex Opaca Ait.
AUA000039362 St. Clair 23 Jul 1967 Thompson, Olivia C.
AUA
Ilex Opaca Ait.
AUA000031874 Chambers 24 Nov 1975 Wilkins, David
270
AUA
Ilex Opaca Ait.
JSU119622 St. Clair 12 Jul 1969 Ellard, Jerry
s.n.
JSU
Ilex opaca Aiton
JSU119627 Tuscaloosa 19 Aug 1975 Whetstone, R. David
6721
JSU
Ilex opaca Aiton
AUA000032182 Monroe 26 Apr 1986 Diamond, Alvin R.
2420
AUA
Ilex Opaca Ait.
UWAL0010716 Pike 05 Dec 2011 Diamond, Alvin R.
22737
UWAL
Ilex opaca Aiton
UNA00023831 Lawrence 26 Jun 1989 Harding, Dennis
s.n.
UNA
Ilex opaca Ait.
TROY000030594 Geneva 28 Jan 2010 Diamond, Alvin R.
21133
TROY
Ilex opaca Aiton
JSU119405 Cherokee 15 Apr 1967 Sanford, S.T.
s.n.
JSU
Ilex opaca Aiton
AMAL00005819 Crenshaw 17 Feb 2002 Diamond, Alvin R.
12911
AMAL
Ilex opaca Ait.
UWAL0018912 Autauga 08 Jan 2015 Diamond, Alvin R.
25824
UWAL
Ilex opaca Aiton
UNA00020692 Tuscaloosa s.d. Smith, Eugene A.
s.n.
UNA
Ilex opaca Ait.
TROY000001185 Pike 01 Apr 1999 Martin, Brian H.
45
TROY
Ilex opaca Aiton
JSU119630 Jackson 5 Jul 1969 Bible, Rebecca
s.n.
JSU
Ilex opaca Aiton
AMAL00020040 Monroe 01 Mar 2011 Barger, T. Wayne
RH-24
AMAL
Ilex opaca Ait.
UWAL0000308 Monroe 20 Sep 2007 Diamond, Alvin R.
18297
UWAL
Ilex opaca Aiton
TROY000001176 Escambia 04 Feb 1995 Brittain, Cynthia L.
1
TROY
Ilex opaca Aiton
BRIT73400 Cleburne 24 Aug 1997 Ballard, J. Mark
7399
BRIT
Ilex opaca Aiton
BRIT73407 Crenshaw 17 Feb 2002 Diamond, Alvin R.
12911
BRIT
Ilex opaca Aiton
JSU119629 Winston 3 Aug 1975 Whetstone, R. David
5712
JSU
Ilex opaca Aiton
AUA000032186 Pike 3 May 1988 Diamond, Alvin R.
4636
AUA
Ilex Opaca Ait.
UNA00053950 Blount 19 Aug 1997 Keener, Brian R.
344
UNA
Ilex opaca Ait.
BRIT73425 Mobile 15 Jul 1969 Kral, Robert
35659
BRIT
Ilex opaca Aiton
JSU119414 Cleburne 24 Aug 1997 Ballard, J. Mark
7399
JSU
Ilex opaca Aiton
AMAL00005817 Calhoun 13 May 1988 Snowden, K.E.
327
AMAL
Ilex opaca Aiton
AUA000032192 Wilcox 7 Dec 2005 Diamond, Alvin R.
16083
AUA
Ilex Opaca Ait.
UWAL0030375 Bibb 10 Jun 2015 Davenport, L.J.
5879
UWAL
Ilex opaca Aiton
TROY000051615 Tallapoosa 01 Oct 2020 Diamond, Alvin R.
29475
TROY
Ilex opaca Aiton
UNAF0000513 Franklin 05 Oct 1989 Pounders, Jeff
s.n.
UNAF
Ilex opaca Aiton
JSU119625 Talladega 24 Jan 1993 Dasinger, Adam P.
32
JSU
Ilex opaca Aiton
JSU119395 Blount 15 May 1993 Whetstone, R. David
s.n.
JSU
Ilex opaca Aiton
AUA000031876 Choctaw 23 Apr 1994 Crouch, Virginia E.
878
AUA
Ilex Opaca Ait.
UWAL0003031 Washington 16 Apr 2011 Keener, Brian R.
6364
UWAL
Ilex opaca Ait.
UNA00020708 Lawrence 22 Aug 1949 Segars, Charles B.
183
UNA
Ilex opaca Ait.
TROY000035052 Bullock 22 Mar 2012 Dykes, Bailee
379
TROY
Ilex opaca Aiton
JSU119393 Baldwin 1 Sep 1997 Brodeur, Kristin R.
087
JSU
Ilex opaca Aiton
UWAL0050587 Sumter 07 Apr 2021 Ryan, Emma D.
79
UWAL
Ilex opaca Aiton
TROY000032712 Covington 20 Dec 2010 Diamond, Alvin R.
21722
TROY
Ilex opaca Aiton
BRIT73422 Lee 21 Apr 1970 Blankenchip, B.P.
38
BRIT
Ilex opaca Aiton
BRIT73418 Henry 20 Jan 2002 Diamond, Alvin R.
12867
BRIT
Ilex opaca Aiton
JSU119408 Clarke 31 Oct 1997 Whetstone, R. David
18988
JSU
Ilex opaca Aiton
AUA000032179 Monroe 3 Jul 1972 Hybart, Michael G.
119
AUA
Ilex Opaca Ait.
TROY000001169 Conecuh 05 Jan 2002 Diamond, Alvin R.
12849
TROY
Ilex opaca Aiton
BRIT73404 Conecuh 16 Nov 1978 Kral, Robert
63174
BRIT
Ilex opaca Aiton
JSU119412 Cleburne 27 Jul 1995 Hruska, Susan E.
849
JSU
Ilex opaca Aiton
AUA000031877 Cleburne 11 Oct 1956 Adams, W.H.
AUA
Ilex Opaca Ait.
AUA000031886 Dale 9 May 1991 Diamond, Alvin R.
7417
AUA
Ilex Opaca Ait.
UWAL0035745 Sumter 02 Oct 1993 Stanton, Lee E.
B-012
UWAL
Ilex opaca
TROY000042063 Houston 02 Apr 2015 Diamond, Alvin R.
26115
TROY
Ilex opaca Aiton
UNAF0000528 Winston 07 Jul 1990 Riechenberg, Chris
s.n.
UNAF
Ilex opaca Aiton
JSU119396 Blount 5 May 1993 Dasinger, Adam P.
77
JSU
Ilex opaca Aiton
AMAL00018568 Monroe 20 Sep 2007 Diamond, Alvin R.
18297
AMAL
Ilex opaca Ait.
AUA000031907 Lee 23 Apr 1949 Cunningham, Hugh B.
AUA
Ilex Opaca Ait.
UWAL0012927 Washington 27 Jul 2012 England, J. Kevin
3810
UWAL
Ilex opaca Aiton
UNA00020706 Washington 24 Jul 1949 Segars, Charles B.
193
UNA
Ilex opaca Ait.
TROY000014269 Marion 03 Jun 2002 Keener, Brian R.
2163
TROY
Ilex opaca Aiton
JSU119409 Clarke 24 Jun 1984 Whetstone, R. David
14310
JSU
Ilex opaca Aiton
UWAL0019645 Blount 07 Mar 1991 Kral, Robert
78341
UWAL
Ilex opaca Ait.
TROY000001178 Geneva 26 Jan 2002 Diamond, Alvin R.
12880
TROY
Ilex opaca Aiton
BRIT73426 Pike 12 Jun 1957 Quarterman, E.
157
BRIT
Ilex opaca Aiton
AUA000031879 Coffee 19 Jul 1989 Diamond, Alvin R.
6010
AUA
Ilex Opaca Ait.
UNA00056410 Perry 02 Aug 1995 Birk, Michelle
369
UNA
Ilex opaca Ait.
TROY000001179 Henry 20 Jan 2002 Diamond, Alvin R.
12867
TROY
Ilex opaca Aiton
BRIT73423 Lee 11 Apr 1969 Chitwood, John C.
s.n.
BRIT
Ilex opaca Aiton
BRIT73416 Geneva 26 Jan 2002 Diamond, Alvin R.
12880
BRIT
Ilex opaca Aiton
AUA000031865 Baldwin 10 Oct 2007 Barger, T. Wayne
PERD-94
AUA
Ilex Opaca Ait.
AUA000032191 Washington 16 Apr 2011 Keener, Brian R.
6364
AUA
Ilex Opaca Ait.
TROY000001166 Barbour 05 Jan 2006 Diamond, Alvin R.
16112
TROY
Ilex opaca Aiton
BRIT73403 Conecuh 18 Sep 1993 Kral, Robert
83283
BRIT
Ilex opaca Aiton
JSU119413 Cleburne 4 May 1995 Hruska, Susan E.
688
JSU
Ilex opaca Aiton
AUA000031900 Lee Anonymous
AUA
Ilex Opaca Ait.
AUA000031913 Macon 20 Jul 1972 Luckett, Larry
123
AUA
Ilex Opaca Ait.
UNA00050251 Randolph March 1986 Nixon, C.F.
2081
UNA
Ilex opaca Ait.
TROY000036228 Montgomery 02 Dec 2012 Diamond, Alvin R.
23579
TROY
Ilex opaca Aiton
UNAF0000522 Madison 29 Sep 1989 Wieseman, Kevin
s.n.
UNAF
Ilex opaca Aiton
JSU119623 St. Clair 12 Jul 1969 Day, Randy
s.n.
JSU
Ilex opaca Aiton
AMAL00001921 Monroe 24 Oct 2006 Diamond, Alvin R.
17148
AMAL
Ilex opaca Ait.
AUA000031866 Baldwin 25 Jul 1969 Ellis, H.C.
073
AUA
Ilex Opaca Ait.
NY01343410 s.d. Samuel B. Buckley
s.n.
NY
Ilex opaca Aiton
UNA00020693 Tuscaloosa _ Apr _ Smith, Eugene A.
s.n.
UNA
Ilex opaca Ait.
UNA00057606 Escambia 10 Feb 2002 Diamond, Alvin R.
12897
UNA
Ilex opaca Ait.
TROY000001182 Montgomery 13 Apr 1991 Funk, H.
1
TROY
Ilex opaca Aiton
BRIT73402 Conecuh 05 Jan 2002 Diamond, Alvin R.
12849
BRIT
Ilex opaca Aiton
BRIT73414 Escambia 29 Dec 1985 Diamond, Alvin R.
2035
BRIT
Ilex opaca Aiton
SAMF0005106 Tuscaloosa 24 Apr 1962 May, Meredith
78
SAMF
Ilex opaca Aiton
AUA000042035 Bullock 21 Jun 2016 Barger, T. Wayne
WLE-79
AUA
Ilex Opaca Ait.
UWAL0013552 Montgomery 02 Dec 2012 Diamond, Alvin R.
23579
UWAL
Ilex opaca Aiton
TROY000001171 Crenshaw 26 Sep 1999 Diamond, Alvin R.
11608
TROY
Ilex opaca Aiton
BRIT73427 Washington 21 Sep 1969 Kral, Robert
37290
BRIT
Ilex opaca Aiton
JSU119431 Marshall 16 Oct 1994 Spaulding, Daniel D.
6791
JSU
Ilex opaca Aiton
AUA000031901 Lee 5 Jun 1960 Baker, Shelby
AUA
Ilex Opaca Ait.
AUA000031888 Dale 21 Apr 1979 Merrill, L.E.
070
AUA
Ilex Opaca Ait.
UNA00057613 Geneva 26 Jan 2002 Diamond, Alvin R.
12880
UNA
Ilex opaca Ait.
TROY000001168 Coffee 08 Dec 1999 Martin, Brian H.
394
TROY
Ilex opaca Aiton
USAM000001433 28 Apr 1969 Erwin, Francene
5
USAM
Ilex opaca Aiton
JSU119438 Randolph 19 Aug 1986 Nixon, Christopher F.
2790
JSU
Ilex opaca Aiton
AMAL00005816 Butler 29 Dec 2000 Diamond, Alvin R.
12260
AMAL
Ilex opaca Ait.
AUA000031905 Lee 30 May 1960 Holdaway, Joy
AUA
Ilex Opaca Ait.
UNA00020690 Bibb 10 Oct 1982 Burckhalter, Robert
53
UNA
Ilex opaca Ait.
TROY000041877 Autauga 08 Jan 2015 Diamond, Alvin R.
25824
TROY
Ilex opaca Aiton
TROY000051652 Lowndes 30 Jan 2021 Diamond, Alvin R.
29540
TROY
Ilex opaca Aiton
BRIT73396 Choctaw 12 Apr 1994 Crouch, Virginia
878
BRIT
Ilex opaca Aiton
BRIT73391 Bibb 27 Jul 2003 Diamond, Alvin R.
14042
BRIT
Ilex opaca Aiton
SAMF0005105 Marion 15 May 1954 McCullough, Herbert A.
s.n.
SAMF
Ilex opaca Aiton
AUA000040240 Houston 21 Jan 2015 Barger, T. Wayne
DTP-19
AUA
Ilex Opaca Ait.
UWAL0011311 Bullock 22 Mar 2012 Dykes, Bailee
379
UWAL
Ilex opaca Aiton
TROY000001172 Crenshaw 17 Feb 2002 Diamond, Alvin R.
12911
TROY
Ilex opaca Aiton
BRIT73390 Baldwin 21 Oct 1969 Kral, Robert
38160
BRIT
Ilex opaca Aiton
JSU146260 Cleburne 13 Apr 1996 Hutchinson, Francine
2376
JSU
Ilex opaca Aiton
AUA000031867 Baldwin 26 Apr 1969 Atkinson, Jr., A.W.
096
AUA
Ilex Opaca Ait.
AUA000032187 Pike 1999 Martin, Brian
45
AUA
Ilex Opaca Ait.
UNA00050466 Bibb 28 Sep 1995 Birk, Michelle
518
UNA
Ilex opaca Ait.
TROY000001184 Pike 22 Dec 2000 Diamond, Alvin R.
12250
TROY
Ilex opaca Aiton
USAM000001425 Montgomery 20 Apr 1969 Andrews, Romona
13
USAM
Ilex opaca Aiton var. opaca
JSU119403 Cherokee 11 Jun 1987 Dickson, Catherine C.
404
JSU
Ilex opaca Aiton
AMAL00005823 Geneva 26 Jan 2002 Diamond, Alvin R.
12880
AMAL
Ilex opaca Ait.
AUA000031890 DeKalb 25 Jul 1970 Holcombe, W.H.
086
AUA
Ilex Opaca Ait.
UNA00020702 Clarke 08 May 1987 Burckhalter, Robert
983
UNA
Ilex opaca Ait.
TROY000014268 Tallapoosa 18 Apr 2006 Barger, T. Wayne
78
TROY
Ilex opaca Aiton
TROY000014265 Cherokee 08 Jul 1969 Payne, Jan E.
n.s.
TROY
Ilex opaca Aiton
UNA00065870 Conecuh 16 Nov 1978 Kral, Robert
63174
UNA
Ilex opaca Ait.
TROY000001175 Dale 20 May 1998 Prazinko, Brian F.
s.n.
TROY
Ilex opaca Aiton
BRIT73412 Elmore 13 Jan 2002 Diamond, Alvin R.
12856
BRIT
Ilex opaca Aiton
JSU119411 Clay 5 Oct 1992 Ballard, J. Mark
680
JSU
Ilex opaca Aiton
AUA000032180 Monroe 29 Dec 2005 Diamond, Alvin R.
16110
AUA
Ilex Opaca Ait.
UWAL0031819 Conecuh 18 Sep 1993 Kral, Robert
83283
UWAL
Ilex opaca Aiton
TROY000029358 Marengo 24 Sep 2008 England, J. Kevin
1180
TROY
Ilex opaca Aiton
JSU119424 Elmore 27 Apr 1979 Whetstone, R. David
9640
JSU
Ilex opaca Aiton
AUA000031897 Houston 6 Aug 1993 Diamond, Alvin R.
8734
AUA
Ilex Opaca Ait.
AUA000031915 Macon 21 Jul 1972 Moody, Gary
094
AUA
Ilex Opaca Ait.
UNA00056342 Dale 26 Dec 1998 MacDonald, John R.
12477
UNA
Ilex opaca Ait.
BRIT73392 Blount 07 Mar 1991 Kral, Robert
78344
BRIT
Ilex opaca Aiton
SAMF0005109 Bibb 01 May 2008 Davenport, Lawrence J.
3814
SAMF
Ilex opaca Aiton
JSU119626 Tallapoosa 2 Feb 1992 Clearman, F.W., Jr.
013
JSU
Ilex opaca Aiton
JSU119439 Randolph 27 Aug 1986 Nixon, Christopher F.
2895
JSU
Ilex opaca Aiton
AMAL00010371 Cherokee 10 Sep 1993 Dobson, Terri L.
191
AMAL
Ilex opaca Aiton
UWAL0013102 Elmore 12 Mar 2013 Diamond, Alvin R.
23793
UWAL
Ilex opaca Aiton
UNA00073799 Washington 27 Jul 2012 England, J. Kevin
3810
UNA
Ilex opaca Ait.
TROY000039049 Elmore 09 Jan 2014 Diamond, Alvin R.
24729
TROY
Ilex opaca Aiton
BRIT73419 Henry 07 Apr 1997 MacDonald, John R.
10194
BRIT
Ilex opaca Aiton
UNA00057621 Henry 20 Jan 2002 Diamond, Alvin R.
12867
UNA
Ilex opaca Ait.
TROY000001174 Dale 09 Nov 1999 Pennington, Tiffany
297
TROY
Ilex opaca Aiton
BRIT73413 Escambia 10 Feb 2002 Diamond, Alvin R.
12897
BRIT
Ilex opaca Aiton
SAMF0005107 Lawrence 04 May 1957 Westbrook, John
s.n.
SAMF
Ilex opaca Aiton
AUA000031893 Escambia 29 Dec 1985 Diamond, Alvin R.
2036
AUA
Ilex Opaca Ait.
UWAL0000305 Escambia 09 Apr 2009 Diamond, Alvin R.
20562
UWAL
Ilex opaca Aiton
TROY000014266 Bibb 27 Jul 2003 Diamond, Alvin R.
14042
TROY
Ilex opaca Aiton
JSU119624 Talladega 26 May 1993 Whetstone, R. David
16752
JSU
Ilex opaca Aiton
AUA000031868 Barbour 5 Jan 2006 Diamond, Alvin R.
16112
AUA
Ilex Opaca Ait.
AUA000031919 Mobile 22 Nov 1975 Mills, S.F.
177
AUA
Ilex Opaca Ait.
UNA00057689 Cherokee 24 Sep 1987 Dickson, Catherine C.
564
UNA
Ilex opaca Ait.
BRIT73397 Clarke 06 Jun 1971 Kral, Robert
43038
BRIT
Ilex opaca Aiton
USAM000001432 Mobile 30 May 1968 LeLong, Michel G.
4683
USAM
Ilex opaca Aiton
JSU119436 Randolph 24 Jun 1986 Nixon, Christopher F.
2207
JSU
Ilex opaca Aiton
AMAL00016035 Monroe 29 Dec 2005 Diamond, Alvin R.
16110
AMAL
Ilex opaca Ait.
AUA000031903 Lee 12 Aug 1970 Hopson, George R.
00162
AUA
Ilex Opaca Ait.
UNA00065362 Winston 04 Mar 2005 England, J. Kevin
269
UNA
Ilex opaca Ait.
TROY000024650 Escambia 09 Apr 2009 Diamond, Alvin R.
20562
TROY
Ilex opaca Aiton
BRIT73410 DeKalb 27 Jul 1977 Kral, Robert
60634
BRIT
Ilex opaca Aiton
JSU143077 Cherokee 08 Jul 1969 Mostowski, Howard S.
s.n.
JSU
Ilex opaca Aiton
BRIT73395 Chilton 23 Mar 1969 Kral, Robert
33949a
BRIT
Ilex opaca Aiton
JSU119399 Calhoun 30 Apr 1969 Morello, Sandra
s.n.
JSU
Ilex opaca Aiton
AUA000032183 Monroe 2 Sep 1986 Diamond, Alvin R.
2788
AUA
Ilex Opaca Ait.
UWAL0038350 Monroe 27 Apr 1975 LeLong, Michel G.
8424
UWAL
Ilex opaca Aiton
BRIT73417 Geneva 26 Oct 1996 McDaniel, Sidney
7963
BRIT
Ilex opaca Aiton
UNAF0000515 Lauderdale -- Jun 1980 Chandler, Jane
s.n.
UNAF
Ilex opaca Aiton
UNAF0000527 Winston 02 Dec 1989 Hatton, Jonathan
s.n.
UNAF
Ilex opaca Aiton
JSU119410 Clay 7 Jul 1999 Brodeur, Kristin R.
2044
JSU
Ilex opaca Aiton
AUA000031870 Bibb 4 May 1979 Beckett, Scott W.
137
AUA
Ilex Opaca Ait.
AUA000032181 Monroe 3 Jul 1981 Tucker, S.
14
AUA
Ilex Opaca Ait.
UWAL0038344 Monroe 27 Apr 1975 LeLong, Michel G.
8442
UWAL
Ilex opaca Aiton
JSU119401 Calhoun 12 Sep 1972 Ginn, Larry
s.n.
JSU
Ilex opaca Aiton
JSU119422 Cullman 20 Jun 1975 Whetstone, R. David
4765
JSU
Ilex opaca Aiton
AUA000031881 Conecuh 26 Apr 1986 Diamond, Alvin R.
2436
AUA
Ilex Opaca Ait.
UWAL0015849 Pike 03 Jan 2014 Diamond, Alvin R.
24713
UWAL
Ilex opaca Aiton
UNA00023873 Lawrence 26 Jun 1989 Harding, Dennis
s.n.
UNA
Ilex opaca Ait.
TROY000001167 Butler 29 Dec 2000 Diamond, Alvin R.
12260
TROY
Ilex opaca Aiton
JSU119425 Etowah 3 Jul 1969 Smith, Doris
s.n.
JSU
Ilex opaca Aiton
AUA000031872 Bullock 11 Sep 1987 Diamond, Alvin R.
3575
AUA
Ilex Opaca Ait.
UWAL0016203 Elmore 09 Jan 2014 Diamond, Alvin R.
24729
UWAL
Ilex opaca Aiton
SAMF0008368 DeKalb 31 Apr 2011 Boatwright, J.L.
37
SAMF
Ilex opaca Aiton
TROY000001173 Dale 10 Apr 1997 Rundell, Hannelore
65
TROY
Ilex opaca Aiton
JSU119416 Cleburne 15 Oct 1972 L'Eplattenier, R.
s.n.
JSU
Ilex opaca Aiton
AMAL00014739 Tallapoosa 18 Apr 2006 Barger, T. Wayne
78
AMAL
Ilex opaca Aiton
UWAL0048171 Monroe 16 Jan 2017 Diamond, Alvin R.
27759
UWAL
Ilex opaca Aiton
JSU119440 Randolph 1 Mar 1986 Nixon, Christopher F.
2081
JSU
Ilex opaca Aiton
AUA000031887 Dale 23 Jun 1991 Diamond, Alvin R.
7485
AUA
Ilex Opaca Ait.
TROY000001177 Escambia 10 Feb 2002 Diamond, Alvin R.
12897
TROY
Ilex opaca Aiton
BRIT73401 Cleburne 09 Apr 1968 Payton, Charles
s.n.
BRIT
Ilex opaca Aiton
JSU119432 Marshall 19 Jul 1982 Minor, Donna K.
s.n.
JSU
Ilex opaca Aiton
AUA000031895 Escambia 2 Jul 1988 Diamond, Alvin R.
5143
AUA
Ilex Opaca Ait.
UWAL0039561 Autauga 11 Nov 2016 Parker, Matthew R.
302
UWAL
Ilex opaca Aiton
BRIT73409 DeKalb 19 May 1987 McDaniel, Sidney
29369
BRIT
Ilex opaca Aiton
JSU119428 Jackson 2 Apr 1999 Brodeur, Kristin R.
1675
JSU
Ilex opaca Aiton
AMAL00017853 Conecuh 16 Nov 1978 Kral, Robert
63174
AMAL
Ilex opaca Ait.
UWAL0035931 Blount 21 May 2013 Anderson, Kelly
159
UWAL
Ilex opaca Aiton
BRIT73420 Henry 10 Jun 1998 MacDonald, John R.
11437
BRIT
Ilex opaca Aiton
UNA00020691 Mobile 15 May 1895 Mohr, Charles
s.n.
UNA
Ilex opaca Ait.
BRIT73393 Blount 27 Jun 1962 Avery
s.n.
BRIT
Ilex opaca Aiton
BRIT73405 Conecuh 16 Dec 1985 Diamond, Alvin R.
1961
BRIT
Ilex opaca Aiton
JSU119430 Marshall 26 Feb 1992 Spaulding, Daniel D.
1089
JSU
Ilex opaca Aiton
AUA000031885 Dale 29 Nov 1991 Diamond, Alvin R.
7858
AUA
Ilex Opaca Ait.
UNA00054209 Lamar 08 Apr 2001 Haynes, Robert R.
10066
UNA
Ilex opaca Ait.
TROY000043617 Washington 27 Jun 2012 England, J. Kevin
3810
TROY
Ilex opaca Aiton
JSU119429 Jackson 28 Jul 1975 Whetstone, R. David
5453
JSU
Ilex opaca Aiton
AUA000031892 Escambia 6 Sep 1986 Diamond, Alvin R.
2872
AUA
Ilex Opaca Ait.
AUA000031914 Macon 21 Jul 1972 Young, Becky
100
AUA
Ilex Opaca Ait.
UNA00020688 Hale 25 Sep 1965 Williams, Norris H.
124
UNA
Ilex opaca Ait.
BRIT73399 Clarke 13 Jul 1969 Kral, Robert
35485
BRIT
Ilex opaca Aiton
UNAF0000524 Marion 21 Jun 1980 Musgrove, Mary Beth
s.n.
UNAF
Ilex opaca Aiton
JSU119406 Cherokee 15 Apr 1967 Collier, Gary
s.n.
JSU
Ilex opaca Aiton
JSU119435 Randolph 29 May 1985 Nixon, Christopher F.
1106
JSU
Ilex opaca Aiton
AUA000031911 Macon 23 May 1971 Carroll, Iii, A.B.
110
AUA
Ilex Opaca Ait.
UWAL0002766 Butler 16 Jan 2011 Diamond, Alvin R.
21732
UWAL
Ilex opaca Aiton
UNA00050745 Henry 10 Jun 1998 MacDonald, John R.
11437
UNA
Ilex opaca Ait.
TROY000014399 Monroe 20 Sep 2007 Diamond, Alvin R.
18297
TROY
Ilex opaca Aiton
JSU119394 Bibb 1 Apr 1999 Brodeur, Kristin R.
1627
JSU
Ilex opaca Aiton
JSU119437 Randolph 11 Oct 1985 Nixon, Christopher F.
2017
JSU
Ilex opaca Aiton
UWAL0000818 Geneva 28 Jan 2010 Diamond, Alvin R.
21133
UWAL
Ilex opaca Aiton
TROY000001180 Monroe 24 Oct 2006 Diamond, Alvin R.
17148
TROY
Ilex opaca Aiton
BRIT73394 Calhoun 26 Mar 1968 Payton, Charles
s.n.
BRIT
Ilex opaca Aiton
AUA000032194 Wilcox 2 Sep 1986 Diamond, Alvin R.
2848
AUA
Ilex Opaca Ait.
UWAL0047234 Autauga 14 Dec 2017 Parker, Matthew R.
1486
UWAL
Ilex opaca Aiton
BRIT73428 Blount 01 Jul 1957 Morton, J.T. [Mrs.]
122
BRIT
Ilex opaca Aiton
TROY000001170 Conecuh 16 Nov 1978 Kral, Robert
63174
TROY
Ilex opaca Aiton
JSU119434 Monroe 26 Aug 1972 McCool, Jeff
s.n.
JSU
Ilex opaca Aiton
AMAL00005820 DeKalb 27 Jul 1977 Kral, Robert
60634
AMAL
Ilex opaca Ait.
UWAL0044380 Baldwin 27 Jun 1995 Newton, Troy
3
UWAL
Ilex opaca Aiton
BRIT73408 Dale 26 Dec 1998 MacDonald, John R.
12477
BRIT
Ilex opaca Aiton
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