Galium aparine

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Distribution Map: Based on vouchered plant specimens only. View county names by placing the mouse cursor over a particular county.

Species Distribution Map
Family:RUBIACEAE
Species:Galium aparine L.
Common Name:Cleavers; Sticky Willy; Catchweed Bedstraw
Habitat:Various sites including woodlands, their edges, roadsides and other disturbed sites.
Associated Ecological Communities:**
Growth Habit:Vine
Duration:Annual
Category:Vascular
USDA Symbol:**
Plant Notes:Cleavers is a native or introduced herbaceous annual in the Madder family (Rubiaceae). Some authorities consider it native only to Europe, northern Africa, and Asia, and introduced in North America. It is possible that the species was introduced on the fur and clothing of people and animals crossing the Bering Land Bridge into North America from Asia during the last Ice Age as well as by colonists arriving from Europe in the 1600’s and 1700’s. In Alabama it can be found throughout the state. Cleavers occurs on roadsides, in forest edges and clearing, along streams, and in other disturbed sites. It is an annual with a tap root. The stems are prostrate or trailing over other vegetation and often form mats. The stems are green in color, angled, with short, stiff, backwards pointing teeth along their margins. The leaves occur in whorls of 6-8 per node. The leaves are sessile, linear to oblanceolate in outline, pubescent, with backwards pointing bristles along their margins. Flowers are produced in open, branched, terminal or axillary cymes. Individual flowers have 4 white petals. The fruit is a dry, globular nutlet with hooked bristles that occurs in pairs. Aparine is derived from Greek and means ‘clinging’ or ‘seizing’.—A. Diamond
Taxonomic Notes:**
Status:Native, FACU (NWI), G5 (Global Rank)
References:**

** Not applicable or data not available.

Classification

FamilyRUBIACEAE - Madder/Coffee family
Genus Galium
Species Galium aparine L. - Cleavers; Sticky Willy; Catchweed Bedstraw

Citation

Citation Galium aparine Linnaeus, Sp. Pl. 1: 108. 1753.
Basionym: **
Type: Without data (lectotype: LINN 129.3). Lectotypified by Jafri, in Jafri & El-Gadi, Fl. Libya 65: 15. 1979.

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Galium aparine var. intermedium Galium aparine Linnaeus, var. intermedium Bonnet, Petite Fl. Paris 191. 1883.Galium intermedium Merat 1831, non Schultes 1809. 
Galium aparine var. microphyllum Galium aparine Linnaeus, var. microphyllum Clos, in Gay, Fl. Chil. 3: 190. 1848. CHILE: Coquimbo: Without data, Gay 68 (holotype: P).
Galium aparine var. minor Galium aparine Linnaeus, var. minor Hooker, Fl. Bor.-Amer. 1: 290. 1834. CANADA: British Columbia: Near Fort Vancouver, s.d., Douglas s.n. (holotype: K?).
Galium aparine var. pseudoaparine Galium aparine Linnaeus, var. pseudoaparine (Grisebach) Spegazzini, Revista Agron. Univ. Nac. La Plata 3: 526. 1897.Galium pseudoaparine Grisebach 1854. 
Galium aparine var. subglabrum Galium aparine Linnaeus, var. subglabrum Petermann, Fl. Lips. Excurs. 122. 1838. GERMANY
Galium aparine var. verum Galium aparine Linnaeus, var. verum Wimmer & Grabowski, Fl. Siles. 1(1): 119. 1827, nom. inadmiss.Galium aparine Linnaeus 1753. 
Galium aparine var. vulgare Galium aparine Linnaeus, var. vulgare Reichenbach, Icon. Fl. Germ. Helv. 17: 94. 1854, nom. inadmiss.Galium aparine Linnaeus 1753. 
Galium australe Galium australe Reiche, Fl. Chile 3: 149. 1900, nom. illegit.Galium chilense Hooker f. 1846. 
Galium chilense Galium chilense Hooker f., Fl. Antarct. 302bis. 1846. CHILE: Asien: Chonos Archipelago, s.d., Darwin s.n. (holotype: K).
Galium chonosense Galium chonosense Clos, in Gay, Fl. Chil. 3: 189. 1848, nom. illegit. et non Hooker f. 1846.Galium chilense Hooker f. 1846. 
Galium hispidum Galium hispidum Willdenow, Enum. Pl. 154. 1809, non Roth 1789, nec Gaertner 1791. EGYPT
Galium intermedium Galium intermedium Merat, Nouv. Fl. Env. Paris, ed. 3. 2: 255. 1831, non Schultes 1809. FRANCE
Galium lappaceum Galium lappaceum Salisbury, Prodr. Stirp. Chap. Allerton 59. 1796, nom. illegit.Galium aparine Linnaeus 1753. 
Galium larecajense Galium larecajense Wernham, J. Bot. 50: 244. 1912. BOLIVIA: La Paz: Prov. Larecaja, vicinity of Sorata, between Munaypata and Challasuyo River, s.d., Mandon 339 (holotype: BM; isotype: NY).
Galium parviflorum Galium parviflorum Maximowicz, Prim. Fl. Amur. 479. 1859, non Moench 1794, nec Rafinesque 1808, nec D. Don 1825.  
Galium pauciflorum Galium pauciflorum Bunge, Enum. Pl. China Bor. 35. 1833, non Willdenow ex de Candolle 1830.  
Galium pseudoaparine Galium pseudoaparine Grisebach, Abh. Königl. Ges. Wiss. Göttingen 6: 125. 1854. CHILE: Magallanes: Near Punta Arenas, s.d., Lechler 1207 (holotype: GOET; isotypes: BR, G, K, P).
Galium scaberrimum Galium scaberrimum Vahl ex Hornemann, Hort. Bot. Hafn. 135. 1813.Galium hispidum Willdenow 1809, non Roth 1789, nec Gaertner 1791. 
Galium sororium Galium sororium Hance, in Walpers, Ann. Bot. Syst. 2: 734. 1852.  
Galium uncinatum Galium uncinatum Gray, Nat. Arr. Brit. Pl. 2: 484. 1821, nom. illegit.Galium aparine Linnaeus 1753. 
Galium aparine var. echinospermum    

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Blount, Bullock, Butler, Calhoun, Chambers, Cherokee, Chilton, Choctaw, Clarke, Clay, Cleburne, Coffee, Colbert, Conecuh, Coosa, Covington, Crenshaw, Cullman, Dale, Dallas, DeKalb, Elmore, Escambia, Etowah, Fayette, Geneva, Greene, Hale, Henry, Houston, Jackson, Jefferson, Lamar, Lauderdale, Lawrence, Lee, Limestone, Lowndes, Madison, Marengo, Marshall, Mobile, Monroe, Montgomery, Morgan, Pickens, Pike, Randolph, Russell, Shelby, St. Clair, Sumter, Tallapoosa, Tuscaloosa, Walker, Washington, Wilcox, Winston

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AUA, JSU, SAMF, TROY, UNA, USAM, UWAL, UWFP

Range of years during which specimens were collected: 1955 - 2000

Barcode / Accession No. County Coll. Date Collector &
Collection No.		 Herbarium &
Herbarium Name Used		 Image
AUA_ACC_48841 Monroe 2 Apr 1988 Diamond, Alvin R.
4369		 AUA
Galium Aparine L.
JSU131857 Cleburne 26 Apr 1995 Hruska, Susan E.
668		 JSU
Galium aparine Linnaeus
UWAL0036100 Blount 19 Apr 2015 Anderson, Kelly
779		 UWAL
Galium aparine L.
UNA00066662 Butler 22 Mar 2007 Diamond, Alvin R.
17331		 UNA
Galium aparine L.
TROY000008652 Dale 20 Feb 2000 Prazinko, Brian F.
s.n.		 TROY
Galium aparine Linnaeus
UWAL0018147 Limestone 25 Apr 2015 England, J. Kevin
5409		 UWAL
Galium aparine Linnaeus
JSU131926 Jackson 20 May 1974 r, da
2713		 JSU
Galium aparine Linnaeus
TROY000053710 Barbour 24 Mar 2022 Diamond, Alvin R.
29838		 TROY
Galium aparine Linnaeus
AUA_ACC_45814 Houston 9 Apr 1971 Gwaltney, John R.
065		 AUA
Galium Aparine L.
UWAL0003890 Covington 18 Mar 2008 Diamond, Alvin R.
18742		 UWAL
Galium aparine L.
JSU131851 Calhoun 11 May 1982 Whetstone, R. David
11025		 JSU
Galium aparine Linnaeus
AUA_ACC_63885 Montgomery 22 Apr 2004 Diamond, Alvin R.
14347		 AUA
Galium Aparine L.
JSU131929 Lauderdale 24 Apr 1969 Henson, Jim
s.n.		 JSU
Galium aparine Linnaeus
TROY000023273 Houston 10 Apr 2008 Diamond, Alvin R.
18915		 TROY
Galium aparine Linnaeus
UWAL0003891 Jackson 12 Apr 2008 Keener, Brian R.
4222		 UWAL
Galium aparine L.
JSU131942 Shelby 4 Apr 1993 Whetstone, R. David
16257		 JSU
Galium aparine Linnaeus
AUA_ACC_73097 Limestone 25 Apr 2015 England, J. Kevin
5409		 AUA
Galium Aparine L.
UNA00054153 Blount 11 Apr 1998 Keener, Brian R.
581		 UNA
Galium aparine L.
AUA_ACC_4357 Lee 25 Apr 1949 Cunningham, Hugh B.
 AUA
Galium Aparine L.
UWAL0003896 Montgomery 30 Apr 2009 Diamond, Alvin R.
20644		 UWAL
Galium aparine L.
UNA00061301 Lawrence 20 Apr 2000 England, J. Kevin
133		 UNA
Galium aparineáL.
TROY000043670 Limestone 25 Apr 2015 England, J. Kevin
5409		 TROY
Galium aparine Linnaeus
UWAL0003894 Marengo 01 May 1993 Harris, J.W.
s.n.		 UWAL
Galium aparine L.
UNA00043516 Mobile 27 Mar 1965 Deramus, Rebecca
D286		 UNA
Galium aparine L.
AUA_ACC_72616 Baldwin 1 Apr 2008 Barger, T. Wayne
PERD-315		 AUA
Galium Aparine L.
JSU131931 Lee 8 Apr 1964 Bozarth, Gene A.
s.n.		 JSU
Galium aparine Linnaeus
UNA00043515 Dallas 11 Apr 1969 Thomas, Joab L.
1944		 UNA
Galium aparine L.
UWAL0033653 Clarke 13 Mar 2016 Keener, Brian R.
9356		 UWAL
Galium aparine Linnaeus
UNA00043520 Dallas 09 Apr 1982 Gunn, Scott C.
396		 UNA
Galium aparine L.
AUA_ACC_27424 Randolph 13 Apr 1975 Rutland, David W.
272		 AUA
Galium Aparine L.
UWAL0037204 Cullman 08 Apr 2016 Spaulding, Daniel D.
14484		 UWAL
Galium aparine L.
UNA00043522 Lawrence 09 May 1987 Haynes, Robert R.
9114		 UNA
Galium aparine L.
AUA_ACC_34763 Pike 30 Mar 1974 Jenkins, J.D.
053		 AUA
Galium Aparine L.
JSU131936 Marshall 13 Mar 1992 Spaulding, Daniel D.
1175		 JSU
Galium aparine Linnaeus
TROY000008648 Butler 22 Mar 2007 Diamond, Alvin R.
17331		 TROY
Galium aparine Linnaeus
AUA_ACC_14619 Lee 8 May 1969 Barnes, W.K.
097		 AUA
Galium Aparine L.
UWAL0050830 Greene 12 Apr 2021 Ryan, Emma D.
94		 UWAL
Galium aparine L.
UNA00043518 Tuscaloosa 17 Apr 1963 Deramus, Rebecca
392		 UNA
Galium aparine L.
UWAL0017645 Houston 02 Apr 2015 Diamond, Alvin R.
26087		 UWAL
Galium aparine Linnaeus
TROY000046251 Elmore 20 Mar 2018 Diamond, Alvin R.
28107		 TROY
Galium aparine Linnaeus
AUA_ACC_48713 Montgomery 10 Apr 1988 Diamond, Alvin R.
4400		 AUA
Galium Aparine L.
JSU131927 Jefferson 30 Mar 1985 Barber, J. Patrick
1364		 JSU
Galium aparine Linnaeus
JSU131921 DeKalb 20 Apr 1997 Ballard, J. Mark
7163		 JSU
Galium aparine Linnaeus
TROY000008653 Lowndes 13 Apr 2006 Diamond, Alvin R.
16240		 TROY
Galium aparine Linnaeus
AUA_ACC_41433 Washington 1 Apr 1973 Kelso, Emily
019		 AUA
Galium Aparine L.
JSU131925 Jackson 18 May 1999 Brodeur, Kristin R.
1819		 JSU
Galium aparine Linnaeus
TROY000008650 Conecuh 07 Apr 2001 Diamond, Alvin R.
12348		 TROY
Galium aparine Linnaeus
UNA00043521 Tuscaloosa 27 Mar 1962 Gibbons, Whit
42		 UNA
Galium aparine L.
TROY000008655 Pike 04 Apr 1997 Holmes, Jeanese
271		 TROY
Galium aparine Linnaeus
SAMF0004192 Jefferson 21 Mar 2006 Davenport, Lawrence J.
3410		 SAMF
Galium aparine Linnaeus
AUA_ACC_28738 Tallapoosa 15 Apr 1976 Rutland, David W.
2060		 AUA
Galium Aparine L.
JSU131928 Jefferson 2 Apr 1976 Atkinson, Tim A.
1081		 JSU
Galium aparine Linnaeus
UNA00059101 Pickens 06 Apr 2002 Spaulding, Daniel D.
11414		 UNA
Galium aparine L.
UWAL0011456 Bullock 22 Feb 2012 Dykes, Bailee
303		 UWAL
Galium aparine Linnaeus
JSU131922 Elmore 27 Apr 1979 Whetstone, R. David
9577		 JSU
Galium aparine Linnaeus
TROY000008649 Coffee 03 Mar 2000 Martin, Brian H.
449		 TROY
Galium aparine Linnaeus
AUA_ACC_50401 Houston 9 Feb 1989 Diamond, Alvin R.
5676		 AUA
Galium Aparine L.
JSU131852 Calhoun 18 Apr 1967 Collier, Gary
s.n.		 JSU
Galium aparine Linnaeus
TROY000022718 Marengo 06 Feb 2009 England, J. Kevin
1589		 TROY
Galium aparine Linnaeus
AUA_ACC_45406 Chambers 14 Apr 1979 Carter, G.A.
033		 AUA
Galium Aparine L.
UWAL0003892 Marengo 06 Feb 2009 England, J. Kevin
1589		 UWAL
Galium aparine L.
JSU131923 Etowah 6 May 1996 Hodge-Spaulding, Lesley M.
4458		 JSU
Galium aparine Linnaeus
TROY000023272 Henry 13 Apr 2008 Diamond, Alvin R.
18976		 TROY
Galium aparine Linnaeus
AUA_ACC_10750 Lee 5 Apr 1963 Keyes, John W.
 AUA
Galium Aparine L.
JSU131939 Randolph 18 Apr 1985 Nixon, Christopher F.
1084		 JSU
Galium aparine Linnaeus
TROY000023270 Covington 18 Mar 2008 Diamond, Alvin R.
18742		 TROY
Galium aparine Linnaeus
AUA_ACC_41434 Elmore 17 Apr 1973 Harwell, W.Charles
146		 AUA
Galium Aparine L.
UNA00044683 Randolph 18 Apr 1985 Nixon, C.F.
1084		 UNA
Galium aparine L.
TROY000023350 Pickens 06 Apr 2002 Spaulding, Daniel D.
11141		 TROY
Galium aparine Linnaeus
SAMF0004191 Cherokee 19 Apr 2010 Davenport, Lawrence J.
4238		 SAMF
Galium aparine Linnaeus
AUA_ACC_29494 Cleburne 16 May 1976 Rutland, David W.
2763		 AUA
Galium Aparine L.
JSU131924 Etowah 1 May 1992 Bailey, D.S.
66		 JSU
Galium aparine Linnaeus
TROY000023269 Butler 27 Mar 2008 Diamond, Alvin R.
18824		 TROY
Galium aparine Linnaeus
UWAL0032305 Jackson 08 Apr 2016 England, J. Kevin
6095		 UWAL
Galium aparine Linnaeus
UNA00043531 Hale 15 Apr 1978 Davenport, Lawrence J.
323		 UNA
Galium aparine L.
AUA_ACC_23814 Russell 3 Apr 1971 Donavan, Lois A.
053		 AUA
Galium Aparine L.
JSU131849 Calhoun 22 Feb 1993 Dasinger, Adam P.
46		 JSU
Galium aparine Linnaeus
TROY000030539 Pike 06 Apr 1991 Mansfield, Shawn Lynn
2		 TROY
Galium aparine Linnaeus
AUA_ACC_35216 Covington 27 Apr 1979 Coggins, W.A.
 AUA
Galium Aparine L.
UWAL0003893 Marengo 17 Apr 2009 England, J. Kevin
1941		 UWAL
Galium aparine L.
UNA00048892 Henry 07 Apr 1997 MacDonald, John R.
10144		 UNA
Galium aparine L.
TROY000023352 Tallapoosa 17 Mar 2002 Spaulding, Daniel D.
11116		 TROY
Galium aparine Linnaeus
JSU131932 Madison 16 Apr 1996 Hruska, Susan E.
1030		 JSU
Galium aparine Linnaeus
TROY000023348 Chilton 14 Apr 2005 Diamond, Alvin R.
15140		 TROY
Galium aparine Linnaeus
AUA_ACC_10676 Lee Taylor, T.D.
 AUA
Galium Aparine L.
UNA00043519 Lee _ May 1897 Baker, Charles F.
s.n.		 UNA
Galium aparine L.
TROY000024641 Montgomery 30 Apr 2009 Diamond, Alvin R.
20644		 TROY
Galium aparine Linnaeus
AUA_ACC_11389 Lee 9 Apr 1966 Searcy, James W.
 AUA
Galium Aparine L.
JSU131847 Baldwin 29 Mar 1998 Brodeur, Kristin R.
435		 JSU
Galium aparine Linnaeus
TROY000023351 Pickens 06 Apr 2002 Spaulding, Daniel D.
11151		 TROY
Galium aparine Linnaeus
AUA_ACC_65636 Lowndes 13 Apr 2006 Diamond, Alvin R.
16240		 AUA
Galium Aparine L.
AUA_ACC_28931 Coosa 20 Apr 1976 Rutland, David W.
2286		 AUA
Galium Aparine L.
JSU131933 Madison 1 May 1972 Whetstone, R. David
RDW-1268		 JSU
Galium aparine Linnaeus
TROY000045773 Autauga 04 Apr 2017 Parker, Matthew R.
544		 TROY
Galium aparine Linnaeus
USAM000013940 Mobile 14 Mar 1968 LeLong, Michel G.
4194		 USAM
Galium aparine Linnaeus
UWAL0052032 Madison 11 Apr 2021 England, J. Kevin
11916		 UWAL
Galium aparine L.
UNA00043527 Tuscaloosa 31 Dec 1957 Harper, Roland M.
4385		 UNA
Galium aparine L.
UWAL0003897 St. Clair 07 Apr 2007 Keener, Brian R.
3445		 UWAL
Galium aparine L.
JSU131856 Clay 18 Mar 1995 Spaulding, Daniel D.
8903		 JSU
Galium aparine Linnaeus
AUA_ACC_35228 Butler 4 May 1979 Crawford, Lloyd C.
 AUA
Galium Aparine L.
JSU131938 Morgan 13 Apr 1997 Allison, James (Jim) R.
s.n.		 JSU
Galium aparine Linnaeus
UWAL0039725 Elmore 20 Mar 2018 Diamond, Alvin R.
28107		 UWAL
Galium aparine Linnaeus
TROY000044815 Geneva 23 Mar 2017 Byrd, Cameron
866		 TROY
Galium aparine Linnaeus
AUA_ACC_48661 Pike 31 Mar 1988 Diamond, Alvin R.
4327A		 AUA
Galium Aparine L.
JSU131855 Chilton 1 Apr 1999 Brodeur, Kristin R.
1597		 JSU
Galium aparine Linnaeus
ALNHS00004393 Lee 04 Apr 2016 Taylor, Chris T.
CSP-289		 ALNHS
Galium aparine L.
UNA00043525 Tuscaloosa 22 Feb 1957 Deason, Temd R.
s.n.		 UNA
Galium aparine L.
SAMF0008262 Walker 03 May 2017 Legg, Mary
43		 SAMF
Galium aparine Linnaeus
AUA_ACC_29208 Cleburne 14 May 1976 Rutland, David W.
2675		 AUA
Galium Aparine L.
JSU131850 Calhoun 16 Apr 1974 Atkinson, Tim A.
137		 JSU
Galium aparine Linnaeus
UNA00043524 Tuscaloosa s.d. Smith, Eugene A.
s.n.		 UNA
Galium aparine L.
AUA_ACC_9400 Lee 15 Apr 1960 Jones, Jr., Samuel B.
454		 AUA
Galium Aparine L.
JSU131860 Colbert 24 May 1974 Whetstone, R. David
3190		 JSU
Galium aparine Linnaeus
TROY000035546 Bullock 22 Feb 2012 Dykes, Bailee
303		 TROY
Galium aparine Linnaeus
AUA_ACC_9706 Lee 5 May 1961 Beasley, P.G.
 AUA
Galium Aparine L.
UNA00043533 Marshall 23 Apr 1977 Ross, Larry J.
97		 UNA
Galium aparine L.
TROY000023274 Wilcox 24 Mar 2008 Diamond, Alvin R.
18796		 TROY
Galium aparine Linnaeus
AUA_ACC_13038 Clay 17 Apr 1965 Brown, Jimmy E.
11		 AUA
Galium Aparine L.
UWAL0003895 Morgan 08 Feb 2009 England, J. Kevin
1592		 UWAL
Galium aparine L.
AUA_ACC_39349 Dallas 10 Apr 1982 Gunn, Scott C.
445		 AUA
Galium Aparine L.
JSU131943 Walker 17 Apr 1980 Whetstone, R. David
9506		 JSU
Galium aparine Linnaeus
AUA_ACC_20871 Colbert 23 Apr 1970 Morris, Janice
 AUA
Galium Aparine L.
JSU131941 Walker 17 Apr 1982 Whetstone, R. David
9492		 JSU
Galium aparine Linnaeus
UNA00059122 Tallapoosa 17 Mar 2002 Spaulding, Daniel D.
11116		 UNA
Galium aparine L.
JSU131858 Cleburne 22 Mar 1995 Hruska, Susan E.
576		 JSU
Galium aparine Linnaeus
AUA000037886 Covington 14 Mar 2012 Hansen, Curtis J.
5403		 AUA
Galium Aparine L.
UNA00062976 Montgomery 22 Apr 2004 Diamond, Alvin R.
14347		 UNA
Galium aparine L.
TROY000008651 Dale 17 Mar 2000 Pennington, Tiffany
420		 TROY
Galium aparine Linnaeus
UWAL0013444 Lawrence 27 Apr 2012 England, J. Kevin
3343		 UWAL
Galium aparine Linnaeus
JSU131854 Calhoun 20 Apr 1953 Pendergrass, Suevilla S.
s.n.		 JSU
Galium aparine Linnaeus
TROY000023349 Lamar 06 Apr 2002 Spaulding, Daniel D.
11173		 TROY
Galium aparine Linnaeus
AUA_ACC_39347 Dallas 9 Apr 1982 Gunn, Scott C.
396		 AUA
Galium Aparine L.
AUA_ACC_29294 Tallapoosa 6 Apr 1976 Rutland, David W.
1996		 AUA
Galium Aparine L.
JSU131937 Marshall 1 May 1973 Whetstone, R. David
RDW-1297		 JSU
Galium aparine Linnaeus
AUA_ACC_49163 Crenshaw 22 Apr 1988 Diamond, Alvin R.
4516		 AUA
Galium Aparine L.
UWAL0043140 Baldwin 19 Mar 2000 Larsen, Harry S.
518		 UWAL
Galium aparine L.
ALNHS00000742 Mobile 04 Apr 2010 Barger, T. Wayne
SP#275		 ALNHS
Galium aparine L.
UWAL0003898 Sumter 10 Apr 2010 Keener, Brian R.
5751		 UWAL
Galium aparine L.
JSU131934 Marshall 2 May 1992 Spaulding, Daniel D.
1503		 JSU
Galium aparine Linnaeus
AUA_ACC_61056 Choctaw 23 Apr 1994 Crouch, Virginia E.
882		 AUA
Galium Aparine L.
JSU131853 Calhoun 28 Mar 1994 Upton, D.L.
s.n.		 JSU
Galium aparine Linnaeus
SAMF0004193 Montgomery 01 Apr 1965 Myers, Martha Crystal
s.n.		 SAMF
Galium aparine Linnaeus
UWFP016190 Escambia 18 Apr 2010 James R. Burkhalter
22686		 UWFP
Galium aparine L.
AUA_ACC_21930 Lauderdale 2 May 1970 Murchison, Grover R.
110		 AUA
Galium Aparine L.
JSU131930 Jefferson 2 Apr 1976 Whetstone, R. David
s.n.		 JSU
Galium aparine Linnaeus
UNA00043526 Tuscaloosa 17 Apr 1959 Williamson, Nelda Kathleen
s.n.		 UNA
Galium aparine L.
UNA00043514 Hale 11 Apr 1966 Maginness, Sherrie E.
s.n.		 UNA
Galium aparine L.
AUA_ACC_14077 Calhoun 16 Apr 1968 O'Neal, Judy
 AUA
Galium Aparine L.
UNA00043517 Tuscaloosa 20 Apr 1955 Chermock, Ottilie D.
s.n.		 UNA
Galium aparine L.
TROY000042454 Houston 02 Apr 2015 Diamond, Alvin R.
26087		 TROY
Galium aparine Linnaeus
UNA00065417 Winston 19 Apr 2005 England, J. Kevin
294		 UNA
Galium aparine L.
TROY000023271 Geneva 03 Apr 2008 Diamond, Alvin R.
18835		 TROY
Galium aparine Linnaeus
AUA000040208 Houston 22 Jan 2015 Barger, T. Wayne
DTP-12		 AUA
Galium Aparine L.
JSU131859 Cleburne 25 Mar 1973 Windsor, Susan
s.n.		 JSU
Galium aparine Linnaeus
UWAL0046587 Autauga 04 Apr 2017 Parker, Matthew R.
544		 UWAL
Galium aparine Linnaeus
UNA00043523 Tuscaloosa 02 May 1959 Edwards, Jackie
s.n.		 UNA
Galium aparine L.
UWAL0049364 Bullock 29 Apr 2017 Horne, Howard E.
3826		 UWAL
Galium aparine Linnaeus
JSU131848 Blount 21 Apr 1973 Whetstone, R. David
RDW-1296		 JSU
Galium aparine Linnaeus
AUA_ACC_41448 Cherokee 16 Apr 1972 Hale, A.R.
064		 AUA
Galium Aparine L.
UWAL0009773 Dallas 04 Feb 2012 Keener, Brian R.
6840		 UWAL
Galium aparine L.
UNA00060630 Pike 20 Mar 2003 Diamond, Alvin R.
13799		 UNA
Galium aparine L.
AUA_ACC_34762 Pike 30 Mar 1974 Hutto, T.A.
107		 AUA
Galium Aparine L.
JSU131935 Marshall 13 Apr 1992 Spaulding, Daniel D.
1341		 JSU
Galium aparine Linnaeus
TROY000008654 Montgomery 22 Apr 2004 Diamond, Alvin R.
14347		 TROY
Galium aparine Linnaeus
SAMF0004190 Tuscaloosa 15 Apr 1963 Williams, Norris H.
50		 SAMF
Galium aparine Linnaeus
UWAL0033881 Fayette 30 Mar 2016 Keener, Brian R.
9410		 UWAL
Galium aparine Linnaeus
JSU131940 St. Clair 5 May 1983 Whetstone, R. David
13062		 JSU
Galium aparine Linnaeus
AUA_ACC_21931 Lee 17 Apr 1970 Williams, Ann
059		 AUA
Galium Aparine L.
TROY000008656 Pike 20 Mar 2003 Diamond, Alvin R.
13799		 TROY
Galium aparine Linnaeus
AUA_ACC_48679 Crenshaw 1 Apr 1988 Diamond, Alvin R.
4343		 AUA
Galium Aparine L.
UWAL0037344 Bullock 29 Mar 2016 Soehren, Eric C.
195		 UWAL
Galium aparine L.
UNA00043513 Tuscaloosa 24 Feb 1955 Farmer, Joe A.
s.n.		 UNA
Galium aparine L.
TROY000049509 Lawrence 27 Apr 2012 England, J. Kevin
3343		 TROY
Galium aparine Linnaeus
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