Asplenium resiliens

Family:ASPLENIACEAE
Species:Asplenium resiliens Kunze
Common Name:Black Stem Spleenwort; Little Ebony Spleenwort
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:**
Plant Notes:Black Stem Spleenwort is a native perennial evergreen fern in the Spleenwort family (Aspleniaceae). It can be found in the northern third of Alabama and in the central-south part of the state. Black Stem Spleenwort grows on cliffs, in ravines, around sinkholes, and near caves in small pockets of soil on limestone or basic rocks. It is a perennial with a fibrous root system. The fronds (leaves) are erect, linear to lanceolate in outline, and once pinnate. The leaves are monomorphic (not differing in shape or size between sterile and fertile leaves). The petiole and rachis are black, glabrous, and shiny. There are 20-40 pairs of opposite (or nearly so) pinnae. The pinnae are oblong in outline with entire or slightly crenate margins. Black Stem Spleenwort reproduces by forming spores. The spores are in sporangia that are grouped into clusters known as sori. There are 2-5 pairs of sori on the underside of each fertile pinnae. The sori have an indusium (a flap of tissue along one side that partially covers the sori). Black Stem Spleenwort is apogamous. This is a type of reproduction in which the sporophyte develops directly from the gametophyte without fusion of gametes. It is believed that this is an adaptation to the dry habitats in which this species occurs.--A. Diamond
Taxonomic Notes:**
Status:Native
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyASPLENIACEAE - Spleenwort family
Genus Asplenium
Species Asplenium resiliens Kunze - Black Stem Spleenwort; Little Ebony Spleenwort

Citation

Citation ASPLENIUM RESILIENS Kunze, Linnaea 18: 331. 1844.
Basionym: Asplenium parvulum M. Martens & Galeotti 1842, non Hooker 1840.
Type: MEXICO: Oaxaca: Near Capulalpan and Hacienda del Carmen, s.d., Galeotti 6462 (holotype: BR).

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Asplenium lealii Asplenium lealii Alston, J. Bot. 78: 20. 1940. ARGENTINA: Mendoza: Las Heras, near Ida de la Casa de Piedra, s.d., Ruiz Leal 4762 (holotype: BM).
Asplenium parvulum Asplenium parvulum M. Martens & Galeotti, Mém. Foug. Mexique 15: 60, pl. 15(3). 1842, non Hooker 1840. MEXICO: Oaxaca: Near Capulalpan and Hacienda del Carmen, s.d., Galeotti 6462 (holotype: BR).
Chamaefilix resiliens Chamaefilix resiliens (Kunze) Farwell, Amer. Midl. Naturalist 12: 269. 1931.Asplenium resiliens Kunze 1844; Asplenium parvulum M. Martens & Galeotti 1842, non Hooker 1840. 

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Bibb, Blount, Butler, Calhoun, Cherokee, Cleburne, Colbert, Conecuh, Covington, Crenshaw, DeKalb, Etowah, Franklin, Jackson, Jefferson, Lauderdale, Lawrence, Limestone, Lowndes, Madison, Marshall, Morgan, St. Clair, Talladega, Tuscaloosa, Wilcox

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: AMAL, AUA, JSU, TROY, UNA, UNAF, UWAL

Range of years during which specimens were collected: -

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
UNA00052450 Conecuh 21 Mar 1987 Robert R. Haynes
8836X
UNA
Asplenium resiliens Kunze
UWAL0031307 Lauderdale 30 Jan 2016 England, J. Kevin
5834
UWAL
Asplenium resiliens Kunze
UNA00052458 Jefferson 22 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052460 Colbert 12 Mar 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
UNA00052438 Tuscaloosa 25 Mar 1955 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU104056 Calhoun 20 May 1986 C. L. Lawler
528
JSU
Asplenium resiliens Kunze
UWAL0012002 Franklin 06 Oct 2012 Keener, Brian R.
7586
UWAL
Asplenium resiliens Kunze
JSU104077 Jackson 24 Oct 1992 J. M. Ballard
752
JSU
Asplenium resiliens Kunze
44030AUA_ACC Conecuh 19 SEP 1985 DIAMOND, JR., ALVIN R.
AUA
ASPLENIUM RESILIENS KUNZE
JSU104065 Cleburne 16 Mar 1995 M. A. Lawler
012
JSU
Asplenium resiliens Kunze
UNA00058779 Marshall 25 Mar 2006 Brian R. Keener
2707
UNA
Asplenium resiliens Kunze
UNA00025627 Colbert 5 Sep 1940 James W. Shaw
s.n.
UNA
Asplenium resiliens Kunze
UWAL0031520 Morgan 21 Dec 2015 England, J. Kevin
5808
UWAL
Asplenium resiliens Kunze
JSU104049 Bibb 18 May 1984 R. D. Whetstone
13946
JSU
Asplenium resiliens Kunze
JSU105174 Marshall 30 May 1992 D. D. Spaulding
1975
JSU
Asplenium resiliens Kunze
UWAL0031363 Jackson 03 Jan 2016 England, J. Kevin
5816
UWAL
Asplenium resiliens Kunze
JSU104075 Etowah 14 Jun 1976 T. A. Atkinson
1656
JSU
Asplenium resiliens Kunze
UWAL0001132 St. Clair 26 May 2008 Keener, Brian R.
4490
UWAL
Asplenium resiliens Kunze.
13398AUA_ACC Calhoun 5 MAR 1968 O'NEAL, JUDY
AUA
ASPLENIUM RESILIENS KUNZE
TROY000040907 Lawrence 27 Oct 2012 J. Kevin England
4017
TROY
Asplenium resiliens Kunze
UNA00053168 Etowah 14 Jul 1954 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052445 Covington 28 Aug 1954 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00058258 Blount 2 May 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU105178 Morgan 11 Mar 1984 C. F. Nixon
581
JSU
Asplenium resiliens Kunze
UNA00052449 Jefferson 5 Mar 1953 B. D. Valentine
s.n.
UNA
Asplenium resiliens Kunze
TROY000013235 Marshall 11 Jun 1993 J. M. Ballard
2530
TROY
Asplenium resiliens Kunze
JSU104066 Colbert 25 Dec 1972 Kenneth Landers
s.n.
JSU
Asplenium resiliens Kunze
JSU105179 Talladega 03 Mar 1984 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
UWAL0001131 Marshall 25 Mar 2006 Keener, Brian R.
2707
UWAL
Asplenium resiliens Kunze.
UNA00057818 Blount 17 Aug 1997 J. M. Ballard
7360
UNA
Asplenium resiliens Kunze
JSU105180 Talladega 07 Mar 1973 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
35672AUA_ACC Colbert 21 JUN 1980 SHORT, JOHN W.
1205
AUA
ASPLENIUM RESILIENS KUNZE
JSU105175 Morgan 28 May 1975 R. D. Whetstone
4053
JSU
Asplenium resiliens Kunze
UWAL0034162 DeKalb 06 May 2015 Spaulding, Dan
14260
UWAL
Asplenium resiliens Kunze
JSU104079 Etowah 01 Oct 1976 T. A. Atkinson
1990
JSU
Asplenium resiliens Kunze
UNA00069777 Jackson 16 Apr 2009 T. Wayne Barger
HC-182
UNA
Asplenium resiliens Kunze
TROY000040915 Jackson 22 Sep 2012 J. Kevin England
3954
TROY
Asplenium resiliens Kunze
UNA00052440 Jackson 12 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052446 Morgan 26 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052464 Blount 2 May 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
UNA00053173 Conecuh 9 Aug 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU105169 Madison 11 May 1996 D. D. Spaulding
8554
JSU
Asplenium resiliens Kunze
UNA00052463 Jefferson 5 Mar 1953 B. D. Valentine
s.n.
UNA
Asplenium resiliens Kunze
UWAL0032331 Jackson 08 Apr 2016 England, J. Kevin
6123
UWAL
Asplenium resiliens Kunze
JSU105162 Jackson 29 Apr 1998 K. R. Brodeur
583
JSU
Asplenium resiliens Kunze
JSU104058 Cherokee 01 May 1973 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
TROY000031650 Lowndes 20 May 2010 Alvin R. Diamond
21347
TROY
Asplenium resiliens Kunze
UNA00052462 Bibb _ Jul 1940 Clemens A. Sergott
s.n.
UNA
Asplenium resiliens Kunze
JSU105173 Marshall 07 Aug 1975 R. D. Whetstone
5999
JSU
Asplenium resiliens Kunze
35701AUA_ACC Conecuh 16 JUL 1980 SHORT, JOHN W.
1225
AUA
ASPLENIUM RESILIENS KUNZE
JSU104051 Bibb 25 Sep 1982 R. D. Whetstone
12546
JSU
Asplenium resiliens Kunze
UWAL0034919 Jackson 16 Apr 2009 Barger, T. Wayne
HC-182
UWAL
Asplenium resiliens Kunze
JSU104064 Cleburne 08 Mar 1984 R. O. Goodloe
043
JSU
Asplenium resiliens Kunze
UNA00052453 Marshall 27 May 1980 Russell A. Meigs
649
UNA
Asplenium resiliens Kunze
UNA00052467 DeKalb 11 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052441 Crenshaw 26 Aug 1954 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00053170 Bibb 15 Jan 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
UNA00053172 Conecuh 9 Aug 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU104060 Cherokee 25 Feb 1992 Christy Harris
20
JSU
Asplenium resiliens Kunze
UNA00052454 Blount 22 Feb 1953 B. D. Valentine
s.n.
UNA
Asplenium resiliens Kunze
JSU105166 Jackson 22 May 1998 K. R. Brodeur
733
JSU
Asplenium resiliens Kunze
UWAL0001133 Wilcox 14 May 2009 Keener, Brian R.
5312
UWAL
Asplenium resiliens Kunze.
AMAL00005318 Cherokee 16 Feb 1989 Daniel Spaulding
25
AMAL
Asplenium resiliens Kunze
JSU105164 Jackson 01 Jun 1993 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
TROY000033301 Wilcox 20 May 2010 Alvin R. Diamond
21327
TROY
Asplenium resiliens Kunze
UNA00052459 Lauderdale 14 Mar 1953 Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU105167 Jackson 05 Aug 1975 R. D. Whetstone
5818
JSU
Asplenium resiliens Kunze
32550AUA_ACC Bibb 12 JUL 1977 SHORT, JOHN W.
914
AUA
ASPLENIUM RESILIENS KUNZE
JSU104068 Etowah 22 Mar 1984 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
UWAL0011062 Butler 29 Aug 2012 Webb, Wayne
s.n.
UWAL
Asplenium resiliens Kunze
JSU104062 Cherokee 25 Feb 1992 F. W. Clearman, Jr
026
JSU
Asplenium resiliens Kunze
JSU104057 Calhoun 31 May 1982 R. D. Whetstone
11239
JSU
Asplenium resiliens Kunze
UNA00052466 Jackson 5 Jul 1980 Robert R. Haynes
7983
UNA
Asplenium resiliens Kunze
UNA00052443 Calhoun 1 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052448 Lawrence 25 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052461 Colbert 12 Mar 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
UNA00052439 Conecuh 9 Aug 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU104076 Calhoun 21 Oct 1986 C. L. Lawler
1432
JSU
Asplenium resiliens Kunze
UWAL0036440 Marshall 13 Aug 2017 England, J. Kevin
8418
UWAL
Asplenium resiliens Kunze
JSU105163 Jackson 30 Aug 1997 J. M. Ballard
7477
JSU
Asplenium resiliens Kunze
UNAF0001117 Calhoun 05 Mar 1968 Watkins, Jerry
s.n.
UNAF
Asplenium resiliens Kunze
41339AUA_ACC Conecuh 27 DEC 1985 DIAMOND, JR., ALVIN R.
2021
AUA
ASPLENIUM RESILIENS KUNZE
JSU104054 Calhoun 05 Mar 1968 Martha Stuart
s.n.
JSU
Asplenium resiliens Kunze
UNA00064867 Blount 28 Feb 1998 Brian R. Keener
522
UNA
Asplenium resiliens Kunze
34359AUA_ACC Bibb 29 JUN 1977 SESSLER, ANN H.
1668
AUA
ASPLENIUM RESILIENS KUNZE
44921AUA_ACC Conecuh 1 JAN 1987 DIAMOND, JR., ALVIN R.
2994
AUA
ASPLENIUM RESILIENS KUNZE
JSU105171 Marshall 20 Sep 1986 R. D. Whetstone
15707
JSU
Asplenium resiliens Kunze
JSU105170 Marshall 13 Aug 1975 D. Miller
s.n.
JSU
Asplenium resiliens Kunze
UWAL0035183 Morgan 04 Mar 2017 England, J. Kevin
7182
UWAL
Asplenium resiliens Kunze
JSU104069 Etowah 21 May 1976 T. A. Atkinson
1513
JSU
Asplenium resiliens Kunze
TROY000013233 Marshall 25 Mar 2006 Brian R. Keener
2707
TROY
Asplenium resiliens Kunze
UWAL0002097 Lowndes 20 May 2010 Diamond, A.R.
21347
UWAL
Asplenium resiliens Kunze
UNA00052451 Blount 22 Feb 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00065286 Lawrence 9 Apr 2003 Kevin England
392
UNA
Asplenium resiliens Kunze
UNA00052465 Bibb 24 Apr 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU104070 Etowah 22 Mar 1984 C. F. Nixon
611
JSU
Asplenium resiliens Kunze
UWAL0001129 Lauderdale 13 Apr 1935 Spieth, Alda M.
s.n.
UWAL
Asplenium resiliens Kunze.
JSU104052 Blount 17 Aug 1997 J. M. Ballard
7360
JSU
Asplenium resiliens Kunze
TROY000013232 Jackson 05 Aug 1975 R.D Whetstone
5768
TROY
Asplenium resiliens Kunze
JSU104067 Etowah 23 Apr 1994 L. M. Hodge
1206
JSU
Asplenium resiliens Kunze
UWAL0012779 Lawrence 27 Oct 2012 England, J. Kevin
4017
UWAL
Asplenium resiliens Kunze
UNA00025623 Colbert 14 Sep 1940 James W. Shaw
s.n.
UNA
Asplenium resiliens Kunze
JSU104061 Cherokee 16 Feb 1984 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
32042AUA_ACC Jackson 9 AUG 1974 SHORT, JOHN W.
198
AUA
ASPLENIUM RESILIENS KUNZE
JSU105176 Morgan 29 Mar 1997 R. D. Whetstone
s.n.
JSU
Asplenium resiliens Kunze
JSU105172 Marshall 26 Feb 1992 D. D. Spaulding
1084
JSU
Asplenium resiliens Kunze
UWAL0001130 Lawrence 25 Oct 2008 England, J. Kevin
1450
UWAL
Asplenium resiliens Kunze.
JSU104080 Etowah 14 Jun 1976 T. A. Atkinson
1656
JSU
Asplenium resiliens Kunze
UWAL0001128 Blount 28 Feb 1998 Keener, Brian R.
0522
UWAL
Asplenium resiliens Kunze.
UWAL0001982 Wilcox 20 May 2010 Diamond, A.R.
21327
UWAL
Asplenium resiliens Kunze
UNA00053169 Etowah 14 Jul 1954 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052447 Franklin 4 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052452 Blount 2 May 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU104063 Cherokee 16 Feb 1984 C. F. Nixon
533
JSU
Asplenium resiliens Kunze
53451AUA_ACC Jackson 11 APR 1991 GUNN, SCOTT C.
1935
AUA
ASPLENIUM RESILIENS KUNZE
TROY000013234 Marshall 12 Jun 1993 J. M. Ballard
2611
TROY
Asplenium resiliens Kunze
JSU104048 Etowah 19 Mar 1995 L. M. Hodge
4093
JSU
Asplenium resiliens Kunze
UWAL0012849 Jackson 22 Sep 2012 England, J. Kevin
3954
UWAL
Asplenium resiliens Kunze
UNA00052468 Calhoun 11 Jun 1953 James C. Wilkes
s.n.
UNA
Asplenium resiliens Kunze
JSU104059 Cherokee 11 May 1983 R. D. Whetstone
13127
JSU
Asplenium resiliens Kunze
32043AUA_ACC Limestone 3 JUL 1977 SHORT, JOHN W.
908
AUA
ASPLENIUM RESILIENS KUNZE
JSU105177 Morgan 28 May 1975 R. D. Whetstone
4053
JSU
Asplenium resiliens Kunze
AMAL00014406 St. Clair 26 May 2008 Brian R. Keener
4490
AMAL
Asplenium resiliens Kunze
UWAL0002647 Jackson 18 Feb 2011 England, J. Kevin
2633
UWAL
Asplenium resiliens Kunze
JSU104078 Etowah 10 Aug 1974 T. A. Atkinson
573
JSU
Asplenium resiliens Kunze
UNA00052456 Madison 23 May 1980 Russell A. Meigs
625
UNA
Asplenium resiliens Kunze
UNA00052457 Jackson 1 Mar 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052444 Morgan 25 Jul 1953 Jesse S. White
s.n.
UNA
Asplenium resiliens Kunze
UNA00052469 Lauderdale 14 Mar 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
UNA00053171 Conecuh 9 Aug 1953 R. L. Chermock
s.n.
UNA
Asplenium resiliens Kunze
JSU104071 Etowah 28 Mar 1978 Cheri Gay
16
JSU
Asplenium resiliens Kunze
UWAL0032857 Morgan 13 Feb 2010 England, J. Kevin
2241
UWAL
Asplenium resiliens Kunze
JSU104072 Etowah 27 Jun 1969 Jerry A. Clonts
s.n.
JSU
Asplenium resiliens Kunze
AUA000031001 Colbert 21 Jun 1980 SHORT, JOHN W.
1205
AUA
ASPLENIUM RESILIENS KUNZE
41997AUA_ACC Conecuh 29 DEC 1985 DIAMOND, JR., ALVIN R.
2045
AUA
ASPLENIUM RESILIENS KUNZE
JSU105165 Jackson 25 Sep 1982 Michael A. Ray
54
JSU
Asplenium resiliens Kunze
UNA00027481 Bibb 17 Nov 1882 Charles Mohr
s.n.
UNA
Asplenium resiliens Kunze
AMAL00019813 Cherokee 09 Aug 1982 John W. Short
1278
AMAL
Asplenium resiliens Kunze
32044AUA_ACC Marshall 10 AUG 1974 SHORT, JOHN W.
206
AUA
ASPLENIUM RESILIENS KUNZE
JSU104050 Bibb 22 Jul 1982 R. D. Whetstone
11592
JSU
Asplenium resiliens Kunze
TROY000046029 Butler 07 Nov 2010 Alvin R. Diamond
1704
TROY
Asplenium resiliens Kunze
JSU104055 Calhoun 05 Mar 1968 Dickie Brown
s.n.
JSU
Asplenium resiliens Kunze
JSU104053 Blount 31 Mar 1976 R. D. Whetstone
7893
JSU
Asplenium resiliens Kunze
JSU105161 Jackson 27 May 1974 T. A. Atkinson
27
JSU
Asplenium resiliens Kunze
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